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List of rove beetle (Staphylinidae) species recorded in Britain

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Subfamily Omaliinae

Tribe Anthophagini

ACIDOTA Stephens, 1829 crenata (Fabricius, 1792) cruentata Mannerheim, 1830

ANTHOBIUM Leach, 1819 1 atrocephalum (Gyllenhal, 1827) unicolor (Marsham, 1802)

ANTHOPHAGUS Gravenhorst, 1802 alpinus (Paykull, 1790) caraboides (Linnaeus, 1758)

DELIPHRUM Erichson, 1839 tectum (Paykull, 1789)

EUCNECOSUM Reitter, 1909 2 brachypterum (Gravenhorst, 1802)

GEODROMICUS Redtenbacher, 1857 longipes (Mannerheim, 1830) nigrita (Müller, P.W.J., 1821)

LESTEVA Latreille, 1796 4 hanseni Lohse, 1953 longoelytrata (Goeze, 1777) monticola Kiesenwetter, 1847 pubescens Mannerheim, 1830 punctata Erichson, 1839 5 sicula Erichson, 1840

OLOPHRUM Erichson, 1839 assimile (Paykull, 1800) consimile (Gyllenhal, 1810) fuscum (Gravenhorst, 1806) piceum (Gyllenhal, 1810)

OROCHARES Kraatz, 1857 angustatus (Erichson, 1840) 7

PHILORINUM Kraatz, 1858 sordidum (Stephens, 1834)

PHYLLODREPOIDEA Ganglbauer, 1895 crenata Ganglbauer, 1895

Tribe Eusphalerini

EUSPHALERUM Kraatz, 1858 luteum (Marsham, 1802) minutum (Fabricius, 1792) primulae (Stephens, 1834) sorbi (Gyllenhal, 1810) 8 sorbicola (Kangas, 1941) 9 torquatum (Marsham, 1802)

Tribe Omaliini

ACROLOCHA Thomson, C.G., 1858 10 minuta (Olivier, 1795) sulcula (Stephens, 1834)

ACRULIA Thomson, C.G., 1858 inflata (Gyllenhal, 1813)

HAPALARAEA Thomson, C.G., 1858 pygmaea (Paykull, 1800) 12

HYPOPYCNA Mulsant & Rey, 1880 rufula (Erichson, 1840)

MICRALYMMA Westwood, 1838 marinum (Ström, 1783)

OMALIUM Gravenhorst, 1802 allardi Fairmaire & Brisout, 1859 caesum Gravenhorst, 1806 excavatum Stephens, 1834 exiguum Gyllenhal, 1810 italicum Bernhauer, 1902 laeviusculum Gyllenhal, 1827 laticolle Kraatz, 1858 oxyacanthae Gravenhorst, 1806 riparium Thomson, C.G., 1857 rivulare (Paykull, 1789) 13 rugatum Mulsant & Rey, 1880 rugulipenne Rye, 1864 septentrionis Thomson, C.G., 1856

PHLOEONOMUS Heer, 1839 punctipennis Thomson, C.G., 1867 pusillus (Gravenhorst, 1806)

PHLOEOSTIBA Thomson, C.G., 1858 14 lapponica (Zetterstedt, 1838) plana (Paykull, 1792) 15

PHYLLODREPA Thomson, C.G., 1859 16 devillei Bernhauer, 1902 gracilicornis (Fairmaire & Laboulbène, 1856) heerii (Heer, 1841) 20 ioptera (Stephens, 1832) vilis (Erichson, 1840) floralis (Paykull, 1789) nigra (Gravenhorst, 1806) puberula Bernhauer, 1903 salicis (Gyllenhal, 1810)

XYLODROMUS Heer, 1839 concinnus (Marsham, 1802) 21 depressus (Gravenhorst, 1802) testaceus (Erichson, 1840) 23

XYLOSTIBA Ganglbauer, 1895 24 bosnica (Bernhauer, 1902) 25 monilicornis (Gyllenhal, 1810)

Tribe Hadrognathini

HADROGNATHUS Schaum, 1852 longipalpis (Mulsant & Rey, 1851) 26

Tribe Coryphiini

CORYPHIUM Stephens, 1834 angusticolle Stephens, 1834

EUDECTUS Redtenbacher, 1857 whitei Sharp, 1871

Subfamily Proteininae

Tribe Proteinini

MEGARTHRUS Stephens, 1829 bellevoyei (Saulcy, 1862) denticollis (Beck, 1817) depressus (Paykull, 1789) 28 hemipterus (Illiger, 1794) prosseni Schatzmayr, 1904

METOPSIA Wollaston, 1854 clypeata (Müller, P.W.J., 1821)

PROTEINUS Latreille, 1796 atomarius Erichson, 1840 30 brachypterus (Fabricius, 1792) crenulatus Pandellé, 1867 laevigatus Hochhuth, 1871 ovalis Stephens, 1834

Subfamily Micropeplinae

ARRHENOPEPLUS Koch, 1937 32 tesserula Curtis, 1828

MICROPEPLUS Latreille, 1809 caelatus Erichson, 1839 33 fulvus Erichson, 1840 porcatus (Paykull, 1789) staphylinoides (Marsham, 1802)

Subfamily Pselaphinae

Supertribe EUPLECTITAE LeConte, 1861

Tribe TRICHONYCHINI Reitter, 1882

Subtribe TRICHONYCHINA Reitter, 1882

AMAURONYX Reitter, 1882 maerkelii (Aubé, 1844)

TRICHONYX Chaudoir, 1845 sulcicollis (Reichenbach, 1816)

Subtribe TRIMIINA Bowman, 1934

TRIMIUM Aubé, 1833 brevicorne (Reichenbach, 1816)

Subtribe BIBLOPORINA Park, 1951

BIBLOPORUS Thomson, C.G., 1859 bicolor (Denny, 1825) minutus Raffray, 1914

Subtribe PANAPHANTINA Jeannel, 1950

BIBLOPLECTUS Reitter, 1882 34 ambiguus (Reichenbach, 1816) delhermi Guillebeau, 1888 minutissimus (Aubé, 1833) pusillus (Denny, 1825) spinosus Raffray, 1914 tenebrosus (Reitter, 1880)

Tribe EUPLECTINI LeConte, 1861

Subtribe EUPLECTINA LeConte, 1861

EUPLECTUS Leach, 1817 35 bescidicus Reitter, 1881 bonvouloiri Reitter, 1881 36 brunneus (Grimmer, 1841) 38 decipiens Raffray, 1910 39 duponti Aubé, 1833 infirmus Raffray, 1910 karstenii (Reichenbach, 1816) kirbii Denny, 1825 mutator Fauvel, 1895 nanus (Reichenbach, 1816) piceus Motschulsky, 1835 punctatus Mulsant, 1861 tholini Guillebeau, 1888 41 sanguineus Denny, 1825 signatus (Reichenbach, 1816)

PLECTOPHLOEUS Reitter, 1891 erichsoni (Aubé, 1844) 42 nitidus (Fairmaire, 1857)

Supertribe BATRISITAE Reitter, 1882

Tribe BATRISINI Reitter, 1882

Subtribe BATRISINA Reitter, 1882

BATRISODES Reitter, 1882 adnexus (Hampe, 1863) delaporti (Aubé, 1833) venustus (Reichenbach, 1816)

Supertribe GONIACERITAE Reitter, 1882

Tribe BYTHININI Raffray, 1890

Subtribe BYTHININA Raffray, 1890

BRYAXIS Kugelann, 1794 bulbifer (Reichenbach, 1816) curtisii (Leach, 1817) puncticollis (Denny, 1825)

BYTHINUS Leach, 1817 burrellii Denny, 1825 macropalpus Aubé, 1833

TYCHOBYTHINUS Ganglbauer, 1896 glabratus (Rye, 1870)

Tribe TYCHINI Raffray, 1904

TYCHUS Leach, 1817 niger (Paykull, 1800) striola Guillebeau, 1888 43

Tribe BRACHYGLUTINI Raffray, 1904

Subtribe BRACHYGLUTINA Raffray, 1904

BRACHYGLUTA Thomson, C.G., 1859 fossulata (Reichenbach, 1816) haematica (Reichenbach, 1816) helferi (Schmidt-Göbel, 1836) pandellei (Saulcy, 1876) simplicior Raffray, 1904 44 sinuata (Aubé, 1833) 44 waterhousei (Rye, 1869) 45

FAGNIEZIA Jeannel, 1950 46 impressus (Panzer, 1803)

REICHENBACHIA Leach, 1826 juncorum (Leach, 1817)

RYBAXIS Saulcy, 1876 longicornis (Leach, 1817)

Supertribe PSELAPHITAE Latreille, 1802

Tribe PSELAPHINI Latreille, 1802

PSELAPHAULAX Reitter, 1909 dresdensis (Herbst, 1792)

PSELAPHUS Herbst, 1792 heisei Herbst, 1792

Supertribe CLAVIGERITAE Leach, 1815

Tribe CLAVIGERINI Leach, 1815

CLAVIGER Preyssler, 1790 longicornis Müller, P.W.J., 1818 48 testaceus Preyssler, 1790 Subfamily PHLOEOCHARINAE Erichson, 1839

PHLOEOCHARIS Mannerheim, 1830 subtilissima Mannerheim, 1830 Subfamily TACHYPORINAE MacLeay, 1825

Tribe TACHYPORINI MacLeay, 1825

CILEA Jacquelin du Val, 1856 silphoides (Linnaeus, 1767)

LAMPRINODES Luze, 1901 saginatus (Gravenhorst, 1806)

SEPEDOPHILUS Gistel, 1856 bipunctatus (Gravenhorst, 1802) constans (Fowler, 1888) immaculatus (Stephens, 1832) littoreus (Linnaeus, 1758) lusitanicus Hammond, 1973 marshami (Stephens, 1832) nigripennis (Stephens, 1832) pedicularius (Gravenhorst, 1802) testaceus (Fabricius, 1792) 49

TACHINUS Gravenhorst, 1802 50 bipustulatus (Fabricius, 1792) 52 corticinus Gravenhorst, 1802 elongatus Gyllenhal, 1810 flavolimbatus Pandellé, 1869 humeralis Gravenhorst, 1802 laticollis Gravenhorst, 1802 lignorum (Linnaeus, 1758) marginellus (Fabricius, 1781) pallipes (Gravenhorst, 1806) proximus Kraatz, 1855 rufipennis Gyllenhal, 1810 rufipes (Linnaeus, 1758) 54 scapularis Stephens, 1832 55 subterraneus (Linnaeus, 1758)

TACHYPORUS Gravenhorst, 1802 atriceps Stephens, 1832 chrysomelinus (Linnaeus, 1758) dispar (Paykull, 1789) 56 formosus Matthews, A.H., 1838 hypnorum (Fabricius, 1775) nitidulus (Fabricius, 1781) 57 obtusus (Linnaeus, 1767) pallidus Sharp, 1871 pusillus Gravenhorst, 1806 quadriscopulatus Pandellé, 1869 scitulus Erichson, 1839 solutus Erichson, 1839 tersus Erichson, 1839 transversalis Gravenhorst, 1806

Tribe MYCETOPORINI Thomson, C.G., 1859

BOLITOBIUS Leach, 1819 castaneus (Stephens, 1832) cingulatus (Mannerheim, 1830)

BRYOPHACIS Reitter, 1909 59 crassicornis (Mäklin, 1847) rugipennis (Pandellé, 1869)

BRYOPORUS Kraatz, 1857 cernuus (Gravenhorst, 1806)

ISCHNOSOMA Stephens, 1829 60 longicorne (Mäklin, 1847) splendidum (Gravenhorst, 1806)

LORDITHON Thomson, C.G., 1858 exoletus (Erichson, 1839) lunulatus (Linnaeus, 1761) thoracicus (Fabricius, 1777) trinotatus (Erichson, 1839)

MYCETOPORUS Mannerheim, 1830 angularis Mulsant & Rey, 1853 baudueri Mulsant & Rey, 1875 bimaculatus Boisduval & Lacordaire, 1835 clavicornis (Stephens, 1832) despectus Strand, A., 1969 erichsonanus Fagel, 1965 lepidus (Gravenhorst, 1806) longulus Mannerheim, 1830 monticola Fowler, 1888 nigricollis Stephens, 1835 piceolus Rey, 1883 punctus (Gravenhorst, 1806) rufescens (Stephens, 1832)

PARABOLITOBIUS Li, Zhao & Sakai, 2000 inclinans (Gravenhorst, 1806) 61

Subfamily Trichophyinae

TRICHOPHYA Mannerheim, 1830 pilicornis (Gyllenhal, 1810)

Subfamily Habrocerinae

HABROCERUS Erichson, 1839 capillaricornis (Gravenhorst, 1806)

Subfamily Aleocharinae

Tribe GYMNUSINI Heer, 1839

GYMNUSA Gravenhorst, 1806 brevicollis (Paykull, 1800) variegata Kiesenwetter, 1845

Tribe DEINOPSINI Sharp, 1883

DEINOPSIS Matthews, A.H., 1838 erosa (Stephens, 1832)

Tribe MYLLAENINI Ganglbauer, 1895

MYLLAENA Erichson, 1837 brevicornis (Matthews, A.H., 1838) dubia (Gravenhorst, 1806) elongata (Matthews, A.H., 1838) fowleri Matthews, A., 1883 62 gracilicornis Fairmaire & Brisout, 1859 63 gracilis (Matthews, A.H., 1838) infuscata Kraatz, 1853 intermedia Erichson, 1837 kraatzi Sharp, 1871 masoni Matthews, A., 1883 64 minuta (Gravenhorst, 1806)

Tribe OXYPODINI Thomson, C.G., 1859

Subtribe OXYPODINA Thomson, C.G., 1859

AMAROCHARA Thomson, C.G., 1858 bonnairei (Fauvel, 1865) 65 forticornis (Boisduval & Lacordaire, 1835) umbrosa (Erichson, 1837)

CALODERA Mannerheim, 1830 66 aethiops (Gravenhorst, 1802) nigrita Mannerheim, 1830 protensa Mannerheim, 1830 riparia Erichson, 1837 rubens (Erichson, 1837) 68 rufescens Kraatz, 1856 uliginosa Erichson, 1837

CHILOMORPHA Krása, 1914 longitarsis (Thomson, C.G., 1867)

COUSYA Mulsant & Rey, 1875 69 defecta Mulsant & Rey, 1875 70 nigrata (Fairmaire & Laboulbène, 1856) 71 nitidiventris (Fagel, 1958) 72

CRATARAEA Thomson, C.G., 1858 suturalis (Mannerheim, 1830)

DEXIOGYIA Thomson, C.G., 1858 corticina (Erichson, 1837)

HAPLOGLOSSA Kraatz, 1856 gentilis (Märkel, 1844) marginalis (Gravenhorst, 1806) nidicola (Fairmaire, 1852) picipennis (Gyllenhal, 1827) villosula (Stephens, 1832)

HYGROPORA Kraatz, 1856 cunctans (Erichson, 1837)

ILYOBATES Kraatz, 1856 bennetti Donisthorpe, 1914 nigricollis (Paykull, 1800) propinquus (Aubé, 1850)

ISCHNOGLOSSA Kraatz, 1856 obscura Wunderle, 1990 75 prolixa (Gravenhorst, 1802) turcica Wunderle, 1992 76

MNIUSA Mulsant & Rey, 1875 77 incrassata (Mulsant & Rey, 1852)

OCALEA Erichson, 1837 badia Erichson, 1837 latipennis Sharp, 1870 picata (Stephens, 1832) rivularis Miller, L., 1851

OCYUSA Kraatz, 1856 maura (Erichson, 1837) picina (Aubé, 1850)

OXYPODA Mannerheim, 1831 acuminata (Stephens, 1832) alternans (Gravenhorst, 1802) annularis Mannerheim, 1830 brachyptera (Stephens, 1832) brevicornis (Stephens, 1832) carbonaria (Heer, 1841) elongatula Aubé, 1850 exoleta Erichson, 1839 ferruginea Erichson, 1839 flavicornis Kraatz, 1856 formiceticola Märkel, 1841 haemorrhoa (Mannerheim, 1830) induta Mulsant & Rey, 1861 islandica Kraatz, 1857 lentula Erichson, 1837 longipes Mulsant & Rey, 1861 lurida Wollaston, 1857 mutata Sharp, 1871 81 nigricornis Motschulsky, 1860 nigrocincta Mulsant & Rey, 1875 opaca (Gravenhorst, 1802) praecox Erichson, 1839 82 procerula Mannerheim, 1830 recondita Kraatz, 1856 soror Thomson, C.G., 1855 spectabilis Märkel, 1844 tarda Sharp, 1871 83 tirolensis Gredler, 1863 vittata Märkel, 1842

PAROCYUSA Bernhauer, 1902 longitarsis (Erichson, 1837) rubicunda (Erichson, 1837)

PHLOEOPORA Erichson, 1837 bernhaueri Lohse, 1984 concolor Kraatz, 1856 86 corticalis (Gravenhorst, 1802) nitidiventris Fauvel, 1904 87 testacea (Mannerheim, 1830)

STICHOGLOSSA Fairmaire & Laboulbène, 1856 semirufa (Erichson, 1839)

THIASOPHILA Kraatz, 1856 angulata (Erichson, 1837) inquilina (Märkel, 1842)

Subtribe DINARDINA Mulsant & Rey, 1873

DINARDA Leach, 1819 dentata (Gravenhorst, 1806) hagensi Wasmann, 1889 88 maerkeli Kiesenwetter, 1843 pygmaea Wasmann, 1894 89

HOMOEUSA Kraatz, 1856 acuminata (Märkel, 1842)

Subtribe MEOTICINA Seevers, 1978

MEOTICA Mulsant & Rey, 1873 90 anglica Benick in Muona, 1991 91 exilis (Knoch, 1806) exillima Sharp, 1915 filiformis (Motschulsky, 1860) 92 pallens (Redtenbacher, 1849) 94

Subtribe TACHYUSINA Thomson, C.G., 1859

BRACHYUSA Mulsant & Rey, 1874 concolor (Erichson, 1839)

DACRILA Mulsant & Rey, 1874 95 fallax (Kraatz, 1856) pruinosa (Kraatz, 1856) 96

DASYGNYPETA Lohse, 1974 97 velata (Erichson, 1837) 98

GNYPETA Thomson, C.G., 1858 caerulea (Sahlberg, C.R., 1831) carbonaria (Mannerheim, 1830) ripicola (Kiesenwetter, 1844) 98 rubrior Tottenham, 1939

ISCHNOPODA Stephens, 1835 99 leucopus (Marsham, 1802) umbratica (Erichson, 1837)

TACHYUSA Erichson, 1837 100 coarctata (Erichson, 1837) constricta (Erichson, 1837) objecta (Mulsant & Rey, 1870) 101 scitula (Erichson, 1837)

THINONOMA Thomson, C.G., 1859 99 atra (Gravenhorst, 1806)

Tribe ATHETINI Casey, 1910

Subtribe ATHETINA Casey, 1910

ACROTONA Thomson, C.G., 1859 102 aterrima (Gravenhorst, 1802) benicki (Allen, 1940) 103 exigua (Erichson, 1837) muscorum (Brisout, 1860) obfuscata (Gravenhorst, 1802) parens (Mulsant & Rey, 1852) parvula (Mannerheim, 1830) pseudotenera (Cameron, 1933) 105 pygmaea (Gravenhorst, 1802) sylvicola (Kraatz, 1856) troglodytes (Motschulsky, 1858)

ADOTA Casey, 1909 maritima (Mannerheim, 1843) 108

ALAOBIA Thomson, C.G., 1858 110 gagatina (Baudi, 1848) hybrida (Sharp, 1869) linderi (Brisout, 1863) 111 pallidicornis (Thomson, C.G., 1856) scapularis (Sahlberg, C.R., 1831) sodalis (Erichson, 1837) subglabra (Sharp, 1869) 112 taxiceroides Munster, 1932 113 trinotata (Kraatz, 1856)

ALEVONOTA Thomson, C.G., 1859 aurantiaca Fauvel, 1895 egregia (Rye, 1876) gracilenta (Erichson, 1839) rufotestacea (Kraatz, 1856)

ALIANTA Thomson, C.G., 1858 incana (Erichson, 1837)

ALOCONOTA Thomson, C.G., 1858 cambrica (Wollaston, 1855) currax (Kraatz, 1856) eichhoffi (Scriba, 1867) gregaria (Erichson, 1839) insecta (Thomson, C.G., 1856) mihoki Bernhauer, 1913 114 planifrons (Waterhouse, G.R., 1864) subgrandis Brundin, 1954 116 sulcifrons (Stephens, 1832) coulsoni (Last, 1952) languida (Erichson, 1837) 118 longicollis (Mulsant & Rey, 1852) 119

AMIDOBIA Thomson, C.G., 1858 talpa (Heer, 1841)

AMISCHA Thomson, C.G., 1858 analis (Gravenhorst, 1802) bifoveolata (Mannerheim, 1831) decipiens (Sharp, 1869) forcipata Mulsant & Rey, 1873 nigrofusca (Stephens, 1829)

ANOPLETA Mulsant & Rey, 1874 122 corvina (Thomson, C.G., 1856) kochi (Roubal, 1937) 124 puberula (Sharp, 1869) soedermani (Bernhauer, 1931) 127

ATHETA Thomson, C.G., 1858 aeneicollis (Sharp, 1869) aquatica (Thomson, C.G., 1852) aquatilis (Thomson, C.G., 1867) brunneipennis (Thomson, C.G., 1852) castanoptera (Mannerheim, 1830) ebenina (Mulsant & Rey, 1874) graminicola (Gravenhorst, 1806) heymesi Hubenthal, 1913 129 hypnorum (Kiesenwetter, 1850) incognita (Sharp, 1869) laevicauda Sahlberg, J., 1876 130 triangulum (Kraatz, 1856) xanthopus (Thomson, C.G., 1856) autumnalis (Erichson, 1839) 132 basicornis (Mulsant & Rey, 1852) boletophila (Thomson, C.G., 1856) britanniae (Bernhauer & Scheerpeltz, 1926) crassicornis (Fabricius, 1792) diversa (Sharp, 1869) divisa (Märkel, 1844) euryptera (Stephens, 1832) fungicola (Thomson, C.G., 1852) harwoodi (Williams, 1930) intermedia (Thomson, C.G., 1852) 134 liturata (Stephens, 1832) nidicola (Johansen, 1914) nigritula (Gravenhorst, 1802) 135 oblita (Erichson, 1839) paracrassicornis Brundin, 1954 136 pilicornis (Thomson, C.G., 1852) procera (Kraatz, 1856) 137 ravilla (Erichson, 1839) strandiella (Brundin, 1954) vaga (Heer, 1839)

BADURA Mulsant & Rey, 1873 139 macrocera (Thomson, C.G., 1856) puncticollis (Benick, 1938)

BESSOBIA Thomson, C.G., 1858 140 excellens (Kraatz, 1856) fungivora (Thomson, C.G., 1867) monticola (Thomson, C.G., 1852) occulta (Erichson, 1837)

BOREOPHILIA Benick, 1973 eremita (Rye, 1866) 141

BRUNDINIA Tottenham, 1949 marina (Mulsant & Rey, 1853) 142 meridionalis (Mulsant & Rey, 1853)

CADAVEROTA Yosii & Sawada, 1976 143 cadaverina (Brisout, 1860) hansseni (Strand, 1943) 144

CALLICERUS Gravenhorst, 1802 obscurus Gravenhorst, 1802 rigidicornis (Erichson, 1839)

CERITAXA Mulsant & Rey, 1873 146 dilaticornis (Kraatz, 1856) 147 pervagata (Benick, 1974) 149 testaceipes (Heer, 1839)

CHAETIDA Mulsant & Rey, 1874 150 longicornis (Gravenhorst, 1802)

COPROTHASSA Thomson, C.G., 1859 151 melanaria (Mannerheim, 1830)

DADOBIA Thomson, C.G., 1858 immersa (Erichson, 1837)

DALOTIA Casey, 1910 coriaria (Kraatz, 1856) 152

DATOMICRA Mulsant & Rey, 1874 153 canescens (Sharp, 1869) celata (Erichson, 1837) 154 dadopora (Thomson, C.G., 1867) nigra (Kraatz, 1856) sordidula (Erichson, 1837) 155 zosterae (Thomson, C.G., 1856) 156

DILACRA Thomson, C.G., 1858 157 luteipes (Erichson, 1837) vilis (Erichson, 1837) 158

DIMETROTA Mulsant & Rey, 1873 159 aeneipennis (Thomson, C.G., 1856) atramentaria (Gyllenhal, 1810) cauta (Erichson, 1837) cinnamoptera (Thomson, C.G., 1856) ischnocera (Thomson, C.G., 1870) laevana (Mulsant & Rey, 1852) marcida (Erichson, 1837) nigripes (Thomson, C.G., 1856) setigera (Sharp, 1869)

DINARAEA Thomson, C.G., 1858 aequata (Erichson, 1837) angustula (Gyllenhal, 1810) linearis (Gravenhorst, 1802)

DOCHMONOTA Thomson, C.G., 1859 clancula (Erichson, 1837)

ENALODROMA Thomson, C.G., 1859 hepatica (Erichson, 1839)

GEOSTIBA Thomson, C.G., 1858 162 circellaris (Gravenhorst, 1806)

HALOBRECTA Thomson, C.G., 1858 algae (Hardy, 1851) 163 algophila (Fenyes, 1909) 164 flavipes Thomson, C.G., 1861 princeps (Sharp, 1869) 165

HYDROSMECTA Thomson, C.G., 1858 delicatissima (Bernhauer, 1908) 167 delicatula (Sharp, 1869) eximia (Sharp, 1869) fragilis (Kraatz, 1854) longula (Heer, 1839) subtilissima (Kraatz, 1854)

LIOGLUTA Thomson, C.G., 1858 alpestris (Heer, 1839) granigera (Kiesenwetter, 1850) 171 longiuscula (Gravenhorst, 1802) microptera Thomson, C.G., 1867 pagana (Erichson, 1839)

LYPROCORRHE Thomson, C.G., 1859 anceps (Erichson, 1837)

MICRODOTA Mulsant & Rey, 1873 173 aegra (Heer, 1841) amicula (Stephens, 1832) atomaria (Kraatz, 1856) 174 atricolor (Sharp, 1869) benickiella (Brundin, 1948) boreella (Brundin, 1948) excelsa (Bernhauer, 1911) glabricula (Thomson, C.G., 1867) indubia (Sharp, 1869) inquinula (Gravenhorst, 1802) liliputana (Brisout, 1860) 175 minuscula (Brisout, 1860) palleola (Erichson, 1837) 176 spatuloides (Benick, 1939) subtilis (Scriba, 1866)

MOCYTA Mulsant & Rey, 1874 177 amplicollis (Mulsant & Rey, 1873) clientula (Erichson, 1839) 178 fungi (Gravenhorst, 1806) fussi (Bernhauer, 1908) orbata (Erichson, 1837) orphana (Erichson, 1837)

MYCETOTA Ádám, 1987 179 fimorum (Brisout, 1860) 180 laticollis (Stephens, 1832)

NEHEMITROPIA Lohse, 1971 lividipennis (Mannerheim, 1830)

NEOHILARA Lohse, 1971 subterranea (Mulsant & Rey, 1853)

NOTOTHECTA Thomson, C.G., 1858 confusa (Märkel, 1844) flavipes (Gravenhorst, 1806)

OREOSTIBA Ganglbauer, 1895 182 tibialis (Heer, 1839)

OUSIPALIA des Gozis, 1886 caesula (Erichson, 1839)

PACHNIDA Mulsant & Rey, 1874 nigella (Erichson, 1837)

PACHYATHETA Munster, 1925 183 cribrata (Kraatz, 1856) mortuorum (Thomson, C.G., 1867) 184

PARAMEOTICA Ganglbauer, 1895 185 difficilis (Brisout, 1860)

PARANOPLETA Brundin in Hansen, 1954 inhabilis (Kraatz, 1856)

PHILHYGRA Mulsant & Rey, 1873 186 arctica (Thomson, C.G., 1856) britteni Joy, 1913 debilis (Erichson, 1837) deformis (Kraatz, 1856) elongatula (Gravenhorst, 1802) fallaciosa (Sharp, 1869) gyllenhalii (Thomson, C.G., 1856) hygrobia (Thomson, C.G., 1856) 187 hygrotopora (Kraatz, 1856) luridipennis (Mannerheim, 1830) malleus (Joy, 1913) melanocera (Thomson, C.G., 1856) obtusangula (Joy, 1913) palustris (Kiesenwetter, 1844) parca (Mulsant & Rey, 1873) scotica (Elliman, 1909) terminalis (Gravenhorst, 1806) volans (Scriba, 1859)

PLATARAEA Thomson, C.G., 1858 brunnea (Fabricius, 1798)

PYCNOTA Mulsant & Rey, 1874 paradoxa (Mulsant & Rey, 1861)

RHAGOCNEME Munster, 1923 189 subsinuata (Erichson, 1839)

SCHISTOGLOSSA Kraatz, 1856 aubei (Brisout, 1860) bergvalli Palm, 1968 190 curtipennis (Sharp, 1869) gemina (Erichson, 1837) viduata (Erichson, 1837)

THINOBAENA Thomson, C.G., 1859 150 vestita (Gravenhorst, 1806)

TRAUMOECIA Mulsant & Rey, 1874 192 picipes (Thomson, C.G., 1856)

TRICHIUSA Casey, 1893 immigrata Lohse, 1984 193

XENOTA Mulsant & Rey, 1874 194 myrmecobia (Kraatz, 1856)

Subtribe THAMIARAEINA Fenyes, 1921

THAMIARAEA Thomson, C.G., 1858 cinnamomea (Gravenhorst, 1802) hospita (Märkel, 1844)

Tribe ALEOCHARINI Fleming, 1821

Subtribe ALEOCHARINA Fleming, 1821

ALEOCHARA Gravenhorst, 1802 195 brevipennis Gravenhorst, 1806 curtula (Goeze, 1777) lata Gravenhorst, 1802 intricata Mannerheim, 1830 ruficornis Gravenhorst, 1802 bilineata Gyllenhal, 1810 binotata Kraatz, 1858 196 bipustulata (Linnaeus, 1761) verna Say, 1836 198 obscurella Gravenhorst, 1806 200 phycophila Allen, 1937 201 grisea Kraatz, 1856 punctatella Motschulsky, 1858 spadicea (Erichson, 1837) cuniculorum Kraatz, 1858 discipennis Mulsant & Rey, 1853 fumata Gravenhorst, 1802 funebris Wollaston, 1864 inconspicua Aubé, 1850 kamila Likovský, 1984 204 lanuginosa Gravenhorst, 1802 lygaea Kraatz, 1862 205 maculata Brisout, 1863 moerens Gyllenhal, 1827 moesta Gravenhorst, 1802 sanguinea (Linnaeus, 1758) sparsa Heer, 1839 stichai Likovský, 1965 tristis Gravenhorst, 1806 villosa Mannerheim, 1830

TINOTUS Sharp, 1883 206 morion (Gravenhorst, 1802)

Tribe LOMECHUSINI Fleming, 1821

Subtribe MYRMEDONIINA Thomson, C.G., 1867

DRUSILLA Leach, 1819 canaliculata (Fabricius, 1787)

MYRMOECIA Mulsant & Rey, 1874 plicata (Erichson, 1837) 207

ZYRAS Stephens, 1835 cognatus (Märkel, 1842) collaris (Märkel, 1842) funestus (Gravenhorst, 1806) haworthi Stephens, 1835 humeralis (Gravenhorst, 1802) laticollis (Märkel, 1842) limbatus (Paykull, 1789) lugens (Gravenhorst, 1802)

Subtribe LOMECHUSINA Fleming, 1821

LOMECHUSA Gravenhorst, 1806 emarginata (Paykull, 1789) paradoxa Gravenhorst, 1806

LOMECHUSOIDES Tottenham, 1939 strumosus (Fabricius, 1792)

Tribe FALAGRIINI Mulsant & Rey, 1873

BOHEMIELLINA Machulka, 1941 flavipennis (Cameron, 1920)

BORBOROPORA Kraatz in Kraatz & Fuss, 1862 kraatzii Fuss in Kraatz & Fuss, 1862 208

CORDALIA Jacobs, 1925 obscura (Gravenhorst, 1802)

FALAGRIA Leach, 1819 caesa Erichson, 1837 sulcatula (Gravenhorst, 1806)

FALAGRIOMA Casey, 1906 thoracica (Stephens, 1832)

MYRMECOCEPHALUS MacLeay, 1873 concinnus (Erichson, 1839) 209

MYRMECOPORA Saulcy, 1865 brevipes Butler, 1909 210 oweni Assing, 1997 211 sulcata (Kiesenwetter, 1850) uvida (Erichson, 1840)

Tribe AUTALIINI Thomson, C.G., 1859

AUTALIA Leach, 1819 impressa (Olivier, 1795) longicornis Scheerpeltz, 1947 puncticollis Sharp, 1864 rivularis (Gravenhorst, 1802)

Tribe HOMALOTINI Heer, 1839

Subtribe GYROPHAENINA Kraatz, 1856

AGARICOCHARA Kraatz, 1856 latissima (Stephens, 1832)

BRACHIDA Mulsant & Rey, 1871 exigua (Heer, 1839)

ENCEPHALUS Stephens, 1832 complicans Stephens, 1832

GYROPHAENA Mannerheim, 1830 affinis Mannerheim, 1830 bihamata Thomson, C.G., 1867 congrua Erichson, 1837 fasciata (Marsham, 1802) 213 gentilis Erichson, 1839 hanseni Strand, A., 1946 joyi Wendeler, 1924 joyioides Wüsthoff, 1937 lucidula Erichson, 1837 manca Erichson, 1839 minima Erichson, 1837 munsteri Strand, A., 1935 nana (Paykull, 1800) poweri Crotch, 1867 pseudonana Strand, A., 1939 215 pulchella Heer, 1839 rousi Dvorák, 1966 216 strictula Erichson, 1839 williamsi Strand, A., 1935

Subtribe HOMALOTINA Heer, 1839

ANOMOGNATHUS Solier, 1849 cuspidatus (Erichson, 1839)

CYPHEA Fauvel, 1863 curtula (Erichson, 1837) 217

HOMALOTA Mannerheim, 1830 plana (Gyllenhal, 1810)

Subtribe SILUSINA Fenyes, 1918

SILUSA Erichson, 1837 rubiginosa Erichson, 1837

Subtribe BOLITOCHARINA Thomson, C.G., 1859

BOLITOCHARA Mannerheim, 1830 bella Märkel, 1844 lucida (Gravenhorst, 1802) mulsanti Sharp, 1875 obliqua Erichson, 1837 pulchra (Gravenhorst, 1806)

EURYUSA Erichson, 1837 optabilis Heer, 1839 sinuata Erichson, 1837

LEPTUSA Kraatz, 1856 218 fumida (Erichson, 1839) norvegica Strand, A., 1941 pulchella (Mannerheim, 1830) ruficollis (Erichson, 1839)

HETEROTA Mulsant & Rey, 1873 plumbea (Waterhouse, G.R., 1858)

PSEUDOMICRODOTA Machulka, 1935 paganettii (Bernhauer, 1909) 219

PSEUDOPASILIA Ganglbauer, 1895 testacea (Brisout, 1863) 221

RHOPALOCERINA Reitter, 1909 clavigera (Scriba, 1859)

TACHYUSIDA Mulsant & Rey, 1871 gracilis (Erichson, 1837)

THECTUROTA Casey, 1893 marchii (Dodero, 1922) 222 williamsi (Bernhauer, 1936) 223

Tribe ACTOCHARINI Bernhauer & Schubert, 1911

ACTOCHARIS Sharp, 1870 readingii Sharp, 1870

Tribe PLACUSINI Mulsant & Rey, 1871

PLACUSA Erichson, 1837 complanata Erichson, 1839 224 depressa Mäklin, 1845 pumilio (Gravenhorst, 1802) tachyporoides (Waltl, 1838)

Tribe PHYTOSINI Thomson, C.G., 1867

ARENA Fauvel, 1862 tabida (Kiesenwetter, 1850)

PHYTOSUS Curtis, 1838 balticus Kraatz, 1859 nigriventris (Chevrolat, 1843) spinifer Curtis, 1838

Tribe DIGLOTTINI Jacobson, 1909

DIGLOTTA Champion, G.C., 1899 mersa (Haliday, 1837) sinuaticollis (Mulsant & Rey, 1871) 226

Tribe HYGRONOMINI Thomson, C.G., 1859

HYGRONOMA Erichson, 1837 dimidiata (Gravenhorst, 1806)

Tribe HYPOCYPHTINI Laporte, 1835

CYPHA Leach, 1819 aprilis (Rey, 1882) discoidea (Erichson, 1839) hanseni (Palm, 1949) 228 laeviuscula (Mannerheim, 1830) longicornis (Paykull, 1800) nitida (Palm, 1935) ovulum (Heer, 1839) 229 pulicaria (Erichson, 1839) punctum (Motschulsky, 1857) seminulum (Erichson, 1839) 229 tarsalis (Luze, 1902) 231

HOLOBUS Solier, 1849 apicatus Erichson, 1837 flavicornis (Lacordaire, 1835)

OLIGOTA Mannerheim, 1830 granaria Erichson, 1837 inflata (Mannerheim, 1830) parva Kraatz, 1862 232 picipes (Stephens, 1832) pumilio Kiesenwetter, 1858 punctulata Heer, 1839 pusillima (Gravenhorst, 1806)

Subfamily Scaphidiinae

Tribe SCAPHIINI Achard, 1924

SCAPHIUM Kirby, 1837 immaculatum (Olivier, 1790) 233

Tribe SCAPHIDIINI Latreille, 1807

SCAPHIDIUM Olivier, 1790 quadrimaculatum Olivier, 1790

Tribe SCAPHISOMATINI Casye, 1894

SCAPHISOMA Leach, 1815 agaricinum (Linnaeus, 1758) assimile Erichson, 1845 boleti (Panzer, 1793) Subfamily PIESTINAE Erichson, 1839

SIAGONIUM Kirby, 1815 quadricorne Kirby, 1815

Subfamily Oxytelinae

Tribe DELEASTERINI Reitter, 1909

DELEASTER Erichson, 1839 dichrous (Gravenhorst, 1802)

SYNTOMIUM Curtis, 1828 aeneum (Müller, P.W.J., 1821)

Tribe COPROPHILINI Heer, 1839

COPROPHILUS Latreille, 1829 234 striatulus (Fabricius, 1792)

Tribe THINOBIINI Sharp, 1887

APLODERUS Stephens, 1833 caelatus (Gravenhorst, 1802)

BLEDIUS Leach, 1819 235 subterraneus Erichson, 1839 limicola Tottenham, 1940 236 spectabilis Kraatz, 1857 tricornis (Herbst, 1784) unicornis (Germar, 1825) fergussoni Joy, 1912 subniger Schneider, 1900 bicornis (Germar, 1822) 238 diota Schiödte, 1866 furcatus (Olivier, 1811) annae Sharp, 1911 240 arcticus Sahlberg, J., 1890 atricapillus (Germar, 1825) 241 crassicollis Lacordaire, 1835 defensus Fauvel, 1872 dissimilis Erichson, 1840 erraticus Erichson, 1839 femoralis (Gyllenhal, 1827) filipes Sharp, 1911 242 fuscipes Rye, 1865 gallicus (Gravenhorst, 1806) longulus Erichson, 1839 occidentalis Bondroit, 1907 opacus (Block, 1799) pallipes (Gravenhorst, 1806) 243 praetermissus Williams, 1929 244 terebrans (Schiödte, 1866)

CARPELIMUS Leach, 1819 bilineatus Stephens, 1834 corticinus (Gravenhorst, 1806) despectus (Baudi, 1869) 245 elongatulus (Erichson, 1839) foveolatus (Sahlberg, C.R., 1832) fuliginosus (Gravenhorst, 1802) gracilis (Mannerheim, 1830) halophilus (Kiesenwetter, 1844) impressus (Boisduval & Lacordaire, 1835) lindrothi Palm, 1942 246 manchuricus (Bernhauer, 1938) obesus (Kiesenwetter, 1844) 248 pusillus (Gravenhorst, 1802) rivularis (Motschulsky, 1860) 249 schneideri (Ganglbauer, 1895) similis Smetana, 1967 250 subtilis (Erichson, 1839) zealandicus (Sharp, 1900) 251

MANDA Blackwelder, 1952 mandibularis (Gyllenhal, 1827)

OCHTHEPHILUS Mulsant & Rey, 1856 252 andalusiacus (Fagel, 1957) angustior (Bernhauer, 1943) aureus (Fauvel, 1871) omalinus (Erichson, 1840)

PLANEUSTOMUS Jacquelin du Val, 1857 flavicollis Fauvel, 1871 253 palpalis (Erichson, 1839)

TEROPALPUS Solier, 1849 unicolor (Sharp, 1900) 254

THINOBIUS Kiesenwetter, 1844 bicolor Joy, 1911 255 brevipennis Kiesenwetter, 1850 ciliatus Kiesenwetter, 1844 crinifer Smetana, 1959 257 longipennis (Heer, 1841) 259 major Kraatz, 1857 newberyi Scheerpeltz, 1925 260

THINODROMUS Kraatz, 1857 arcuatus (Stephens, 1834)

Tribe OXYTELINI Fleming, 1821

ANOTYLUS Thomson, C.G., 1859 clypeonitens (Pandellé, 1867) complanatus (Erichson, 1839) fairmairei (Pandellé, 1867) 261 hamatus (Fairmaire & Laboulbène, 1856) insecatus (Gravenhorst, 1806) inustus (Gravenhorst, 1806) maritimus Thomson, C.G., 1861 mutator (Lohse, 1963) nitidulus (Gravenhorst, 1802) rugosus (Fabricius, 1775) 262 saulcyi (Pandellé, 1867) sculpturatus (Gravenhorst, 1806) tetracarinatus (Block, 1799)

OXYTELUS Gravenhorst, 1802 fulvipes Erichson, 1839 laqueatus (Marsham, 1802) migrator Fauvel, 1904 263 piceus (Linnaeus, 1767) sculptus Gravenhorst, 1806

PLATYSTETHUS Mannerheim, 1831 alutaceus Thomson, C.G., 1861 capito Heer, 1839 cornutus (Gravenhorst, 1802) degener Mulsant & Rey, 1878 nitens (Sahlberg, C.R., 1832) nodifrons (Mannerheim, 1830) arenarius (Fourcroy, 1785)

Subfamily Oxyporinae

OXYPORUS Fabricius, 1775 rufus (Linnaeus, 1758)

Subfamily Steninae

STENUS Latreille, 1796 264 aceris Stephens, 1833 fuscicornis Erichson, 1840 geniculatus Gravenhorst, 1806 glacialis Heer, 1839 impressus Germar, 1824 ludyi Fauvel, 1855 267 ochropus Kiesenwetter, 1858 ossium Stephens, 1833 pallipes Gravenhorst, 1802 palustris Erichson, 1839 subaeneus Erichson, 1840 cicindeloides (Schaller, 1783) fornicatus Stephens, 1833 fulvicornis Stephens, 1833 kiesenwetteri Rosenhauer, 1856 latifrons Erichson, 1839 oscillator Rye, 1870 similis (Herbst, 1784) solutus Erichson, 1840 tarsalis Ljungh, 1810 bifoveolatus Gyllenhal, 1827 binotatus Ljungh, 1804 brevipennis Thomson, C.G., 1851 269 butrintensis Smetana, 1959 canescens Rosenhauer, 1856 flavipes Stephens, 1833 nitidiusculus Stephens, 1833 niveus Fauvel, 1865 pallitarsis Stephens, 1833 picipennis Erichson, 1840 picipes Stephens, 1833 pubescens Stephens, 1833 umbratilis Casey, 1884 argus Gravenhorst, 1806 asphaltinus Erichson, 1840 assequens Rey, 1844 ater Mannerheim, 1830 atratulus Erichson, 1839 biguttatus (Linnaeus, 1758) bimaculatus Gyllenhal, 1810 boops Ljungh, 1810 calcaratus Scriba, 1864 270 canaliculatus Gyllenhal, 1827 carbonarius Gyllenhal, 1827 circularis Gravenhorst, 1802 clavicornis (Scopoli, 1763) comma LeConte, 1863 contumax Assing, 1994 271 europaeus Puthz, 1966 fossulatus Erichson, 1840 fuscipes Gravenhorst, 1802 glabellus Thomson, C.G., 1870 272 guttula Müller, P.W.J., 1821 guynemeri Jacquelin du Val, 1850 incanus Erichson, 1839 incrassatus Erichson, 1839 juno (Paykull, 1789) longitarsis Thomson, C.G., 1851 lustrator Erichson, 1839 melanarius Stephens, 1833 melanopus (Marsham, 1802) morio Gravenhorst, 1806 nanus Stephens, 1833 nitens Stephens, 1833 palposus Zetterstedt, 1838 273 proditor Erichson, 1839 providus Erichson, 1839 pusillus Stephens, 1833 subdepressus Mulsant & Rey, 1861 brunnipes Stephens, 1833 crassus Stephens, 1833 formicetorum Mannerheim, 1843 nigritulus Gyllenhal, 1827 opticus Gravenhorst, 1806

DIANOUS Leach, 1819 coerulescens (Gyllenhal, 1810)

Subfamily Euaesthetinae

Tribe EUAESTHETINI Thomson, C.G., 1859

EUAESTHETUS Gravenhorst, 1806 bipunctatus (Ljungh, 1804) laeviusculus Mannerheim, 1844 ruficapillus Lacordaire, 1835 Subfamily PSEUDOPSINAE Ganglbauer, 1895

PSEUDOPSIS Newman, 1834 sulcata Newman, 1834 Subfamily PAEDERINAE Fleming, 1821

British species are distributed among several subtribes of the tribe Paederini, but these subtribes are not listed here pending a modern revision of the subfamily. Generic names and sequence follow Freude et al. (1974) and Silfverberg (1992), except where stated.

Tribe PAEDERINI Fleming, 1821

ACHENIUM Leach, 1819 depressum (Gravenhorst, 1802) humile (Nicolai, 1822)

ASTENUS Dejean, 1833 Subgenus ASTENOGNATHUS Reitter, 1909 procerus (Gravenhorst, 1806) 

    filiformis (Latreille, 1806) non (Fabricius, 1792)

pulchellus (Heer, 1839) 

    melanurus sensu Townsend, 1948 non Küster, 1853

serpentinus (Motschulsky, 1858) 

    procerus sensu auctt. Brit. partim non (Gravenhorst, 1806)
    subditus (Mulsant & Rey, 1878) 277

Subgenus ASTENUS Dejean, 1833 immaculatus Stephens, 1833 lyonessius (Joy, 1908) 

    angustatus sensu auctt. non (Paykull, 1789)
    longelytratus Palm, 1936
    brevelytratus Coiffait, 1960

HYPOMEDON Mulsant & Rey, 1878 278 

    CHLOECHARIS Lynch Arribálzaga, 1884

debilicornis (Wollaston, 1857) 279

LATHROBIUM Gravenhorst, 1802 280 Subgenus PLATYDOMENE Ganglbauer, 1895 angusticolle Boisduval & Lacordaire, 1835 Subgenus TETARTOPEUS Czwalina, 1888 angustatum Boisduval & Lacordaire, 1835 quadratum (Paykull, 1789) rufonitidum Reitter, 1909 

    fennicum Renkonen, 1938 281

terminatum Gravenhorst, 1802 zetterstedti Rye, 1872 

    punctatum Zetterstedt, 1828 non (Fourcroy, 1785)
    foveatum sensu auctt. non Stephens, 1839

Subgenus LATHROBIUM Gravenhorst, 1802 brunnipes (Fabricius, 1792) dilutum Erichson, 1839 elongatum (Linnaeus, 1767) fovulum Stephens, 1833 

    britannicum (Coiffait, 1972)

fulvipenne (Gravenhorst, 1806) geminum Kraatz, 1857 282 

    boreale Hochhuth, 1851
    volgense Hochhuth, 1851

impressum Heer, 1841 

    filiforme Gravenhorst, 1806 non (Fabricius, 1792)

longulum Gravenhorst, 1802 

    longipenne Fairmaire & Laboulbène, 1856

pallidipenne Hichhuth, 1851 

    ripicola Czwalina, 1888 283

pallidum Nordmann, 1837 rufipenne Gyllenhal, 1813

LITHOCHARIS Dejean, 1833 nigriceps Kraatz, 1859 284 ochracea (Gravenhorst, 1802) 285

LOBRATHIUM Mulsant & Rey, 1878 286 

    LATHROBIUM sensu auctt. partim non Gravenhorst, 1802

multipunctum Gravenhorst, 1802

MEDON Stephens, 1833 287 apicalis (Kraatz, 1857) brunneus (Erichson, 1839) castaneus (Gravenhorst, 1802) dilutus (Erichson, 1839) fusculus (Mannerheim, 1830) piceus (Kraatz, 1858) 288 pocofer (Peyron, 1857) ripicola (Kraatz, 1854)

OCHTHEPHILUM Stephens, 1829 

    CRYPTOBIUM Mannerheim, 1830

fracticorne (Paykull, 1800) 

    brevipenne sensu auctt. non Mulsant & Rey, 1861

jacquelini (Boieldieu, 1859)

PAEDERIDUS Mulsant & Rey, 1878 289 

    PAEDERUS sensu auctt. partim non Fabricius, 1775

rubrothoracicus (Goeze, 1777) 290

PAEDERUS Fabricius, 1775 caligatus Erichson, 1840 fuscipes Curtis, 1826 littoralis Gravenhorst, 1802 riparius (Linnaeus, 1758)

PSEUDOMEDON Mulsant & Rey, 1877 291 

    LITHOCHARIS sensu auctt. partim non Dejean, 1833

obscurellus Erichson, 1840 obsoletus (Nordmann, 1837)

RUGILUS Leach, 1819 

    STILICUS Berthold, 1827

erichsoni (Fauvel, 1867) fragilis (Gravenhorst, 1806) 

    angustatus (Fourcroy, 1785) non (Schrank, 1781)
    scutellatus (Motschulsky, 1858)

geniculatus (Erichson, 1839) orbiculatus (Paykull, 1789) rufipes Germar, 1835 similis (Erichson, 1839) subtilis (Erichson, 1840) 292

SCOPAEUS Erichson, 1840 293 gracilis (Sperk, 1835) laevigatus (Gyllenhal, 1827) 

    gracilis sensu auctt. Brit. partim non (Sperk, 1835)

minutus Erichson, 1840 

    sulcicollis sensu Fowler, 1888 non (Stephens, 1833)
    gracilipes Edmonds, 1933

ryei Wollaston, 1872 294 

    minimus sensu auctt. Brit. non Erichson, 1839 295
    rubidus sensu auctt. Brit. non Mulsant & Rey, 1855

sulcicollis (Stephens, 1833) 

    minutus sensu auctt. partim non Erichson, 1840
    cognatus Mulsant & Rey, 1855
    abbreviatus sensu Edmonds, 1931 non Mulsant & Rey, 1855
    pusillus sensu Tottenham, 1949 non Kiesenwetter, 1843

SUNIUS Stephens, 1829 

    HYPOMEDON sensu auctt. non Mulsant & Rey, 1878

bicolor (Olivier, 1795) melanocephalus (Fabricius, 1792) propinquus (Brisout, 1867) Subfamily STAPHYLININAE Latreille, 1802

Tribe STAPHYLININI Latreille, 1802

Subtribe PHILONTHINA Kirby, 1837

BISNIUS Stephens, 1829 296 

    PHILONTHUS sensu auctt. partim non Stephens, 1829

cephalotes (Gravenhorst, 1802) 297 fimetarius (Gravenhorst, 1802) 

    rigidicornis (Gravenhorst, 1802)

nigriventris (Thomson, C.G., 1867) parcus (Sharp, 1874) 298 pseudoparcus (Brunne, 1976) 299 puella (Nordmann, 1837) scoticus (Joy & Tomlin, 1913) 300 sordidus (Gravenhorst, 1802) 

    pachycephalus (Nordmann, 1837)

subuliformis (Gravenhorst, 1802) 

    fuscus (Gravenhorst, 1802) non (Gmelin in Linnaeus, 1790)

CAFIUS Curtis, 1829 cicatricosus (Erichson, 1840) fucicola Curtis, 1830 xantholoma (Gravenhorst, 1806)

ERICHSONIUS Fauvel, 1874 

    ACTOBIUS Fauvel, 1875

cinerascens (Gravenhorst, 1802) signaticornis (Mulsant & Rey, 1852) ytenensis (Sharp, 1913) 301

GABRIUS Stephens, 1829 302 appendiculatus Sharp, 1910 

    subnigritulus sensu auctt. non Joy, 1913 303

astutoides (Strand, A., 1946) 

    astutus sensu auctt. partim non Erichson, 1840

bishopi Sharp, 1910 breviventer (Sperk, 1835) 

    coxalus (Hochhuth, 1871)
    pennatus Sharp, 1910 304

exiguus (Nordmann, 1837) 305 keysianus Sharp, 1910 nigritulus (Gravenhorst, 1802) osseticus (Kolenati, 1846) 

    vernalis (Gravenhorst, 1806) non (Müller, O.F., 1776)

piliger Mulsant & Rey, 1876 

    stipes Sharp, 1910

splendidulus (Gravenhorst, 1802) trossulus (Nordmann, 1837) velox Sharp, 1910 

    primigenius Joy, 1913

GABRONTHUS Tottenham, 1955 thermarum (Aubé, 1850) 306

NEOBISNIUS Ganglbauer, 1895 lathrobioides (Baudi, 1848) 

    cerrutii Gridelli, 1943
    rubripennis Gridelli, 1943

procerulus (Gravenhorst, 1806) prolixus (Erichson, 1840) villosulus (Stephens, 1833)

PHILONTHUS Stephens, 1829 307 

    SPATULONTHUS Tottenham, 1955 308
    PARAGABRIUS Coiffait, 1963 308

addendus Sharp, 1867 albipes (Gravenhorst, 1802) 

    alpinus sensu auctt. Brit. partim non Eppelsheim, 1875 309

atratus (Gravenhorst, 1802) carbonarius (Gravenhorst, 1802) 

    varius (Gyllenhal, 1810)

cognatus Stephens, 1832 

    fuscipennis (Mannerheim, 1830) non (Block, 1799)

concinnus (Gravenhorst, 1802) 

    ochropus (Gravenhorst, 1802)

confinis Strand, A., 1941 310 coprophilus Jarrige, 1949 311 corruscus (Gravenhorst, 1802) 312 corvinus Erichson, 1839 cruentatus (Gmelin in Linnaeus, 1790) debilis (Gravenhorst, 1802) decorus (Gravenhorst, 1802) dimidiatipennis Erichson, 1840 313 discoideus (Gravenhorst, 1802) ebeninus (Gravenhorst, 1802) fumarius (Gravenhorst, 1806) furcifer Renkonen, 1937 314 intermedius (Lacordaire, 1835) jurgans Tottenham, 1937 

    varians sensu auctt. partim non (Paykull, 1789)

laminatus (Creutzer, 1799) lepidus (Gravenhorst, 1802) longicornis Stephens, 1832 mannerheimi Fauvel, 1869 marginatus (Müller, O.F., 1764) micans (Gravenhorst, 1802) micantoides Benick & Lohse, 1956 

    micans sensu auctt. partim non (Gravenhorst, 1802)

nigrita (Gravenhorst, 1806) nitidicollis (Boisduval & Lacordaire, 1835) 

    bimaculatus (Gravenhorst, 1802) non (Schrank, 1798)

parvicornis (Gravenhorst, 1802) 

    agilis (Gravenhorst, 1806) 315

politus (Linnaeus, 1758) 

    aeneus (Rossi, 1790) non (De Geer, 1774)
    temporalis sensu Crowson, 1970 non Mulsant & Rey, 1853

punctus (Gravenhorst, 1802) quisquiliarius (Gyllenhal, 1810) rectangulus Sharp, 1874 316 rotundicollis (Ménétriés, 1832) rubripennis Stephens, 1832 

    fulvipes sensu auctt. Brit. non (Fabricius, 1792)
    varipes Mulsant & Rey, 1861 317

rufipes (Stephens, 1832) 318 

    immundus sensu auctt. non (Gravenhorst, 1806)

sanguinolentus (Gravenhorst, 1802) spinipes Sharp, 1874 319 splendens (Fabricius, 1792) 320 

    niger (Müller, O.F., 1764)

succicola Thomson, C.G., 1860 

    chalceus sensu Ganglbauer, 1895 non Stephens, 1832

tenuicornis Mulsant & Rey, 1853 

    carbonarius sensu (Gyllenhal, 1810) non (Gravenhorst, 1802)

umbratilis (Gravenhorst, 1802) varians (Paykull, 1789) ventralis (Gravenhorst, 1802)

RABIGUS Mulsant & Rey, 1876 

    PHILONTHUS sensu auctt. partim non Stephens, 1829

pullus Nordmann, 1837

REMUS Holme, 1837 321 

    CAFIUS sensu auctt. partim non Curtis, 1829

sericeus Holme, 1837

Subtribe STAPHYLININA Latreille, 1802

CREOPHILUS Leach, 1819 maxillosus (Linnaeus, 1758)

DINOTHENARUS Thomson, C.G., 1858 

    STAPHYLINUS sensu auctt. partim non Linnaeus, 1758
    TRICHODERMA sensu auctt. non Stephens, 1835
    PLATYDRACUS sensu auctt. partim non Thomson, C.G., 1858

pubescens (De Geer, 1774)

EMUS Leach, 1819 hirtus (Linnaeus, 1758)

OCYPUS Leach, 1819 

    GOERIUS Westwood, 1827
    STAPHYLINUS sensu auctt. partim non Linnaeus, 1758

Subgenus MATIDUS Motschulsky, 1860 brunnipes (Fabricius, 1781) 322 nitens (Schrank, 1781) 322 

    nero (Faldermann, 1835) 323
    similis sensu (Fabricius, 1792) non (Herbst, 1784)

Subgenus OCYPUS Leach, 1819 olens (Müller, O.F., 1764) Devil's Coach-horse ophthalmicus (Scopoli, 1763) Subgenus PSEUDOCYPUS Mulsant & Rey, 1876 324 aeneocephalus (De Geer, 1774) 

    cupreus sensu auctt. Brit. partim non (Rossi, 1790)

fortunatarum (Wollaston, 1871) 

    cupreus sensu auctt. Brit. partim non (Rossi, 1790)

fuscatus (Gravenhorst, 1802)

ONTHOLESTES Ganglbauer, 1895 murinus (Linnaeus, 1758) tessellatus (Geoffroy, 1785)

PLATYDRACUS Thomson, C.G., 1858 

    STAPHYLINUS sensu auctt. partim non Linnaeus, 1758

fulvipes (Scopoli, 1763) latebricola (Gravenhorst, 1806) stercorarius (Olivier, 1795)

STAPHYLINUS Linnaeus, 1758 caesareus Cederhjelm, 1798 dimidiaticornis Gemminger, 1851 

    caesareus sensu auctt. Brit. partim non (Cederhjelm, 1798)
    parumtomentosus Stein, 1903

erythropterus Linnaeus, 1758

TASGIUS Stephens, 1829 325 

    STAPHYLINUS sensu auctt. partim non Linnaeus, 1758
    OCYPUS sensu auctt. partim non Leach, 1819
    ALAPSODUS Tottenham, 1939

Subgenus RAYACHEILA Motschulsky, 1845 326 globulifer (Geoffroy, 1785) 

    siculus Stierlin, 1864 non (Aubé, 1842)
    evitendus Tottenham, 1945

melanarius (Heer, 1839) 

    globulifer sensu auctt. non (Geoffroy, 1785)

morsitans (Rossi, 1790) 

    compressus (Marsham, 1802) 327

winkleri Bernhauer, 1906 Subgenus TASGIUS Stephens, 1829 ater (Gravenhorst, 1802) pedator (Gravenhorst, 1802)

Subtribe QUEDIINA Kraatz, 1857

ACYLOPHORUS Nordmann, 1837 glaberrimus (Herbst, 1784)

EURYPORUS Erichson, 1839 picipes (Paykull, 1800)

HETEROTHOPS Stephens, 1829 binotatus (Gravenhorst, 1802) dissimilis (Gravenhorst, 1802) minutus Wollaston, 1860 328 

    dissimilis sensu auctt. partim non (Gravenhorst, 1802)

niger Kraatz, 1868 329 praevius Erichson, 1839

QUEDIUS Stephens, 1829 Subgenus DISTICHALIUS Casey, 1915 cinctus (Paykull, 1790) Subgenus MICROSAURUS Dejean, 1833 330 aetolicus Kraatz, 1858 brevicornis (Thomson, C.G., 1860) brevis Erichson, 1840 cruentus (Olivier, 1795) fulgidus (Fabricius, 1792) 331 

    ?rufitarsis (Marsham, 1802)

assimilis (Nordmann, 1837) invreae Gridelli, 1924 

    ochripennis sensu auctt. Brit. non Ménétriés, 1832
    assecla sensu auctt. non Mulsant & Rey, 1876

lateralis (Gravenhorst, 1802) longicornis Kraatz, 1857 maurus (Sahlberg, C.R., 1830) mesomelinus (Marsham, 1802) microps Gravenhorst, J.L.C., 1847 

    infuscatus sensu Donisthorpe, 1928 non Erichson, 1840

nigrocaeruleus Fauvel, 1874 puncticollis (Thomson, C.G., 1867) 

    othiniensis Johansen, 1907

scitus (Gravenhorst, 1806) 332 

    analis (Fabricius, 1787)

truncicola Fairmaire & Laboulbène, 1856 

    ventralis (Aragona, 1830) non (Gravenhorst, 1802)

xanthopus Erichson, 1839 Subgenus QUEDIONUCHUS Sharp, 1884 plagiatus Mannerheim, 1843 333 

    glaber (Müller, O.F., 1764)
    flavopterus (Geoffroy, 1785)
    laevigatus (Gyllenhal, 1810) non (Marsham, 1802)

Subgenus QUEDIUS Stephens, 1829 balticus Korge, 1960 

    molochinus sensu Last, 1952 non (Gravenhorst, 1806)

curtipennis Bernhauer, 1908 

    fuliginosus sensu Britten, 1944 non (Gravenhorst, 1802)

fuliginosus (Gravenhorst, 1802) 

    subfuliginosus Britten, 1944

levicollis (Brullé, 1832) 

    tristis (Gravenhorst, 1802) non (Fabricius, 1792)

molochinus (Gravenhorst, 1806) simplicifrons Fairmaire, 1861 334 

    hispanicus Bernhauer, 1898
    pallipes sensu Gridelli, 1924 partim non Lucas, 1849
    secundus Last, 1952

Subgenus RAPHIRUS Stephens, 1829 auricomus Kiesenwetter, 1850 boopoides Munster, 1923 

    picipennis sensu auctt. Brit. non Heer, 1839
    scribae sensu auctt. Brit. non Ganglbauer, 1895

boops (Gravenhorst, 1802) 

    asturicus sensu auctt. non Bernhauer, 1918
    arestor Tottenham, 1948
    ?crius Tottenham, 1948

fulvicollis (Stephens, 1833) fumatus (Stephens, 1833) humeralis Stephens, 1832 

    obliteratus Erichson, 1840
    suturalis sensu auctt. Brit. partim non Kiesenwetter, 1845

maurorufus (Gravenhorst, 1806) nemoralis Baudi, 1848 

    humeralis sensu auctt. Brit. partim non Stephens, 1832
    suturalis sensu auctt. Brit. partim non Kiesenwetter, 1845

nigriceps Kraatz, 1857 nitipennis (Stephens, 1833) 

    attenuatus sensu (Gyllenhal, 1810) non (Gravenhorst, 1806)
    hyperboreus sensu auctt. Brit. non Erichson, 1840

persimilis Mulsant & Rey, 1876 

    boops sensu auctt. Brit. partim non (Gravenhorst, 1802)
    aridulus Jansson, 1939 335
    ?corion Tottenham, 1948 336
    mallius Tottenham, 1948

picipes (Mannerheim, 1830) plancus Erichson, 1840 riparius Kellner, 1843 schatzmayri Gridelli, 1922 scintillans (Gravenhorst, 1806) semiaeneus (Stephens, 1833) semiobscurus (Marsham, 1802) 

    rufipes (Gravenhorst, 1802) non (Linnaeus, 1758)

umbrinus Erichson, 1839 

    pseudoumbrinus Lohse, 1961 337

VELLEIUS Leach, 1819 dilatatus (Fabricius, 1787)

Tribe OTHIINI Thomson, C.G., 1859

ATRECUS Jacquelin du Val, 1856 

    BAPTOLINUS Kraatz, 1857

affinis (Paykull, 1789)

OTHIUS Stephens, 1829 angustus Stephens, 1833 

    melanocephalus (Gravenhorst, 1806) non (Geoffroy, 1785)

laeviusculus Stephens, 1833 lapidicola Märkel & Kiesenwetter, 1848 338 punctulatus (Goeze, 1777) subuliformis Stephens, 1833 

    myrmecophilus Kiesenwetter, 1843 339

Tribe XANTHOLININI Erichson, 1839

GAUROPTERUS Thomson, C.G., 1860 fulgidus (Fabricius, 1787) 340

GYROHYPNUS Leach, 1819 angustatus Stephens, 1833 

    scoticus (Joy, 1913) 341

atratus (Heer, 1839) fracticornis (Müller, O.F., 1776) 

    punctulatus sensu auctt. partim non (Paykull, 1789)

punctulatus (Paykull, 1789) 342 

    liebei sensu auctt. non (Scheerpeltz, 1926)

HYPNOGYRA Casey, 1906 343 

    XANTHOLINUS sensu auctt. partim non Dejean, 1821
    PHALACROLINUS Coiffait, 1972

angularis Ganglbauer, 1895 

    glaber (Gravenhorst, 1806) non (Müller, O.F., 1776)

LEPTACINUS Erichson, 1839 batychrus (Gyllenhal, 1827) formicetorum Märkel, 1841 intermedius Donisthorpe, 1936 

    sulcifrons sensu auctt. non (Stephens, 1833)

pusillus (Stephens, 1833) 

    linearis (Gravenhorst, 1802) non (Olivier, 1795)

MEGALINUS Mulsant & Rey, 1877 344 

    XANTHOLINUS sensu auctt. partim non Dejean, 1821

glabratus (Gravenhorst, 1802)

NUDOBIUS Thomson, C.G., 1860 lentus (Gravenhorst, 1806)

PHACOPHALLUS Coiffait, 1956 parumpunctatus (Gyllenhal, 1827) 345 tricolor (Kraatz, 1859) 346 

    pallidipennis sensu auctt. ?non (Motschulsky, 1858) 347

XANTHOLINUS Dejean, 1821 Subgenus ACANTHOPHALLUS Coiffait, 1956 laevigatus Jacobsen, 1849 

    distans sensu auctt. Brit. non Mulsant & Rey, 1853
    cribripennis Fauvel, 1873

Subgenus PURROLINUS Coiffait, 1956 elegans (Olivier, 1795) 348 

    tricolor sensu auctt. partim non (Fabricius, 1787)
    ?meridionalis (Lacordaire, 1835)
    semirufus sensu auctt. non Reitter, 1901
    jarrigei Coiffait, 1956

tricolor (Fabricius, 1787) 349 

    meyeri Drugmand, 1994

Subgenus XANTHOLINUS Dejean, 1821 gallicus Coiffait, 1956 linearis (Olivier, 1795) longiventris Heer, 1839

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