Paoliida

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Paoliida
Temporal range: Carboniferous–Permian
Fossils of Paoliida. A: Zdenekia silesiensis; B: Darekia sanguinea; C, D1, D2: Paoliidae gen et sp. indet.
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Cohort: Polyneoptera
Order: Paoliida
Handlirsch, 1906
Families
Synonyms

Protoptera Rasnitsyn, 1977

Paoliida is an extinct order of winged insects that lived in the late Paleozoic. Historically, both their systematic position and composition were controversial – for instance they had been considered as palaeodictyopterans, as basal Neoptera, or as stem-group of Pterygota – but recent studies have resolved them as the sister group of Dictyoptera.[1][2][3]

Families and genera[edit]

After Prokop et al. (2014), Guan et al. (2016) and Nel & Poschmann (2021):[1][4][5]

References[edit]

  1. ^ a b Jakub Prokop; Wieslaw Krzemiński; Ewa Krzemińska; Thomas Hörnschemeyer; Jan-Michael Ilger; Carsten Brauckmann; Philippe Grandcolas; André Nel (2014). "Late Palaeozoic Paoliida is the sister group of Dictyoptera (Insecta: Neoptera)". Journal of Systematic Palaeontology. 12 (5): 601–622. doi:10.1080/14772019.2013.823468. S2CID 84407734.
  2. ^ Legendre, Frédéric; Nel, André; Svenson, Gavin J.; Robillard, Tony; Pellens, Roseli; Grandcolas, Philippe; Escriva, Hector (22 July 2015). "Phylogeny of Dictyoptera: Dating the Origin of Cockroaches, Praying Mantises and Termites with Molecular Data and Controlled Fossil Evidence". PLOS ONE. 10 (7): e0130127. Bibcode:2015PLoSO..1030127L. doi:10.1371/journal.pone.0130127. PMC 4511787. PMID 26200914.
  3. ^ Prokop, Jakub; Nel, André; Engel, Michael S. (2023). "Diversity, Form, and Postembryonic Development of Paleozoic Insects". Annual Review of Entomology. 68 (1): 401–429. doi:10.1146/annurev-ento-120220-022637.
  4. ^ Guan, Z.; Prokop, J.; Roques, P.; Lapeyrie, J.; Nel, A. (2016). "Revision of the enigmatic family Anthracoptilidae enlightens the evolution of Palaeozoic stem-dictyopterans". Acta Palaeontologica Polonica. 61 (1): 71–87. doi:10.4202/app.00051.2014.
  5. ^ André Nel; Markus J. Poschmann (2021). "Comparison of the recently described early Permian paoliid genus Permomertovia with the 'eoblattid' genera of the families Permulidae and Permotermopsidae". Palaeoentomology. 4 (1): 39–43. doi:10.11646/palaeoentomology.4.1.8.
  6. ^ Nel, A.; Garrouste, R.; Prokop, J. (2015). "The first African Anthracoptilidae (Insecta: Paoliida) near the Permian–Triassic boundary in Kenya". Zootaxa. 3925 (1): 145–150. doi:10.11646/zootaxa.3925.1.10.
  7. ^ Jakub Prokop; Jacek Szwedo; Jean Lapeyrie; Romain Garrouste; André Nel (2015). "New Middle Permian insects from Salagou Formation of the Lodève Basin in southern France (Insecta: Pterygota)". Annales de la Société Entomologique de France. New Series. 51 (1): 14–51. doi:10.1080/00379271.2015.1054645. S2CID 87501340.
  8. ^ Quispe, L.; Roques, P.; Garrouste, R.; Nel, A. (2021). "Carboniferous Blattinopsidae: revision of Klebsiella and new genus and species from Avion (Insecta, Paoliida)" (PDF). Historical Biology. 34 (3): 383–389. doi:10.1080/08912963.2021.1916817. S2CID 234869567.
  9. ^ Nel, A.; Garrouste, R.; Jouault, C. (2023). "New insects (Paoliida, Dictyoptera) from the Carboniferous outcrop of Tante Victoire in Var, France". Alcheringa: An Australasian Journal of Palaeontology. 47 (3): 305–314. doi:10.1080/03115518.2023.2258977.
  10. ^ Santos, A. A.; Hernández-Orúe, A.; Wappler, T.; Peñalver, E.; Diez, J. B.; Nel, A. (2022). "Late Carboniferous insects from the Iberian Peninsula: State of the art and new taxa". Palaeontographica Abteilung A. 326 (1–6): 1–27. doi:10.1127/pala/2022/0135. S2CID 251938467.