2019 in arthropod paleontology

From Wikipedia, the free encyclopedia
List of years in arthropod paleontology
In paleontology
2016
2017
2018
2019
2020
2021
2022
In science
2016
2017
2018
2019
2020
2021
2022
In reptile paleontology
2016
2017
2018
2019
2020
2021
2022
In paleobotany
2016
2017
2018
2019
2020
2021
2022
In paleoentomology
2016
2017
2018
2019
2020
2021
2022
In paleomalacology
2016
2017
2018
2019
2020
2021
2022
In archosaur paleontology
2016
2017
2018
2019
2020
2021
2022
In mammal paleontology
2016
2017
2018
2019
2020
2021
2022
In paleoichthyology
2016
2017
2018
2019
2020
2021
2022

This list of fossil arthropods described in 2019 is a list of new taxa of trilobites, fossil insects, crustaceans, arachnids and other fossil arthropods of every kind that are scheduled to be described during the year 2019, as well as other significant discoveries and events related to arthropod paleontology that are scheduled to occur in the year 2019.

General research[edit]

  • A revision of higher-level arthropod taxonomy is published by Aria (2019), who proposes the name Cenocondyla for the least inclusive group containing both Mandibulata and Chelicerata.[1]
  • A study on the molecular composition and microanatomy of the eyes of 54-million-year-old crane flies from the Fur Formation (Denmark), evaluating its implications for the knowledge of optical systems of other ancient arthropods (especially trilobites), is published by Lindgren et al. (2019).[2]

Arachnids[edit]

Research[edit]

  • Two new specimens of laniatorid harvestmen (an adult specimen assignable to the species Proholoscotolemon nemastomoides and a smaller specimen belonging or related to the genus Proholoscotolemon) are described from the Eocene Baltic amber by Bartel & Dunlop (2019).[3]
  • A review of the phylogenetic hypotheses of scorpion interrelationships, as well as of scorpion palaeontology, is published by Howard et al. (2019), who also provide a list of scorpion fossil calibrations for use in molecular dating.[4]
  • A fossil spider, possibly a juvenile female golden silk orb-weaver, is described from the Lower Eocene Palana Formation of the Gurha opencast lignite mine (western Rajasthan, India) by Patel, Rana & Selden (2019), representing the first fossil spider from India which wasn't found in amber.[5]
  • A review of the fossil record and evolutionary history of spiders, reevaluating the phylogenetic placement of key fossils and aiming to determine whether there has been a major turnover in the spider fauna between the Mesozoic and Cenozoic, is published online by Magalhaes et al. (2019).[6]

New taxa[edit]

Name Novelty Status Authors Age Type locality Country Notes Images

Caestaranea[7]

Gen. et sp. nov

Valid

Selden, Huang & Garwood

Middle–Late Jurassic

Haifanggou Formation

 China

A palpimanoid spider. Genus includes new species C. jurassica.

Colossopantopodus nanus[8]

Sp. nov

Valid

Sabroux et al.

Late Jurassic

Solnhofen Limestone

 Germany

A sea spider.

Cosmobunus sagani[9]

Sp. nov

Valid

Palencia et al.

Miocene (Burdigalian)

Rubielos de Mora Basin

 Spain

A member of Opiliones, a species of Cosmobunus.

Eatoniana crinita[10]

Sp. nov

Valid

Sidorchuk et al.

Eocene

Baltic amber

 Russia
( Kaliningrad Oblast)

A mite belonging to the family Erythraeidae.

Eomesothele[11]

Gen. et sp. nov

Valid

Wunderlich

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to group Mesothelae, assigned to the new family Eomesothelidae. The type species is E. noninclinata.

Eumesosoma abdelmawlai[12]

Sp. nov

Valid

Elsaka, Mitov & Dunlop

Eocene

Baltic amber

Europe (Baltic Sea region)

A harvestman, a species of Eumesosoma.

?Eurycyde golem[8]

Sp. nov

Valid

Sabroux et al.

Late Jurassic

Solnhofen Limestone

 Germany

A sea spider belonging to the family Ascorhynchidae.

Intermesothele[11]

Gen. et sp. nov

Valid

Wunderlich

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to group Mesothelae, possibly belonging to the family Eomesothelidae. The type species is I. pulcher.

Jinjumegops[13]

Gen. et sp. nov

Valid

Park, Nam & Selden

Early Cretaceous (Albian)

Jinju Formation

 South Korea

A spider belonging to the family Lagonomegopidae. The type species is J. dalingwateri.

Koreamegops[13]

Gen. et sp. nov

Valid

Park, Nam & Selden

Early Cretaceous (Albian)

Jinju Formation

 South Korea

A spider belonging to the family Lagonomegopidae. The type species is K. samsiki.

Lutetiana[14]

Gen. et sp. nov

Valid

Selden & Wappler

Eocene (Lutetian)

Messel pit

 Germany

A spider, possibly a member of the family Cybaeidae. The type species is L. neli.

Metagonia esquincacanoi[15]

Sp. nov

García-Villafuerte

Miocene (Aquitanian)

Mexican amber

 Mexico

A species of Metagonia.

Neoliodes andreneli[16]

Sp. nov

Valid

Arillo et al.

Early Cretaceous

Lebanese amber

 Lebanon

A mite belonging to the family Neoliodidae.

Onychopalpus[7]

Gen. et sp. nov

Valid

Selden, Huang & Garwood

Middle–Late Jurassic

Haifanggou Formation

 China

A palpimanoid spider. Genus includes new species O. thomisoides.

Paragilarovia[12]

Gen. et sp. nov

Valid

Elsaka, Mitov & Dunlop

Eocene

Baltic amber

Europe (Baltic Sea region)

A harvestman belonging to the family Nemastomatidae. Genus includes new species P. hochae.

Procaeculus coineaui[17]

Sp. nov

Valid

Porta et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A mite belonging to the family Caeculidae.

Protolophus hoffeinsi[12]

Sp. nov

Valid

Elsaka, Mitov & Dunlop

Eocene

Baltic amber

Europe (Baltic Sea region)

A harvestman, a species of Protolophus.

Scolotydaeus vlaskini[18]

Sp. nov

Valid

Klimov et al.

Late Eocene

Rovno amber

 Ukraine

A mite belonging to the family Paratydeidae.

Sinaranea brevicrus[7]

Sp. nov

Valid

Selden, Huang & Garwood

Middle–Late Jurassic

Haifanggou Formation

 China

A palpimanoid spider.

Tanytydeus pogrebnyaki[18]

Sp. nov

Valid

Klimov et al.

Late Eocene

Rovno amber

 Ukraine

A mite belonging to the family Paratydeidae.

Crustaceans[edit]

Research[edit]

  • A global dataset of exceptionally preserved muscles and muscle attachment scars in fossil malacostracan specimens is compiled by Klompmaker et al. (2019).[19]
  • A three-dimensionally preserved tanaidacean specimen belonging or related to the species Jurapseudes friedericianus is described from the Middle Jurassic (Callovian) Ornatenton Formation (Germany) by Schädel et al. (2019).[20]
  • A study on the evolutionary history of decapod crustaceans is published by Wolfe et al. (2019).[21]
  • A study on the chemical composition of two specimens of fossilized shrimps from the Cretaceous Romualdo and Ipubi formations (Araripe Basin, Brazil), and on its implications for inferring the course of the processes of fossilization of these crustaceans, is published by Barros et al. (2019).[22]
  • A study on the Middle Jurassic palaeoenvironment of La Voulte (France), as indicated by data from exceptionally preserved eyes of the polychelidan lobster Voulteryon parvulus and from epibiontic brachiopods associated with V. parvulus, is published by Audo et al. (2019).[23]
  • A study on the anatomy of the holotype specimen of a putative Paleocene shark Platyacrodus unicus is published by Bogan, Agnolin & Ezcurra (2019), who reinterpret this specimen as a carapace of a small retroplumid crab belonging to the genus Costacopluma.[24]
  • An achelatan larval fossil with an intermediate type of morphology, interpreted as the oldest occurrence of an achelatan lobster larva reported so far, is described from the Toarcian Posidonia Shale (Germany) by Haug, Haug & Schweigert (2019).[25]
  • Redescription of the isopod species Palaega sismondai Ristori (1891) is published by Hyžný, Pasini & Garassino (2019), who transfer this species to the genus Bathynomus.[26]
  • A moult of a giant isopod is described from the early Oligocene Ranzano Formation by Pasini & Garassino (2019), representing the oldest fossil record of the genus Bathynomus from Italy.[27]
  • A study on the Late Devonian ostracods from the Yangdi and Nandong sections in South China, and on their responses to the Frasnian-Famennian event, is published by Song, Huang & Gong (2019).[28]
  • A diverse earliest Triassic ostracod fauna is reported from the Yangou section in South China by Qiu et al. (2019).[29]
  • A revision of ostracods from the Upper Oligocene to Lower Miocene Pirabas Formation (Brazil) is published by Nogueira, Ramos & Hunt (2019).[30]
  • A study on changes of diversity of Neogene fossil marine ostracods from Java, Indonesia is published by Shin et al. (2019).[31]
  • A study on deep-sea benthic ostracod assemblages from the southern Sea of Japan covering the last 2 million years, evaluating their responses to the Mid-Brunhes Event, orbital-scale climatic cycles, and fluctuations of the Tsushima Warm Current, is published by Huang et al. (2019).[32]
  • A three-dimensionally preserved specimen of a late growth stage of the phosphatocopid species Cyclotron angelini is described from the Furongian strata of northern Poland by Olempska et al. (2019).[33]
  • A study on the ecology of clam shrimps from the Anjiagou and Hengdaozi beds of the Lower Cretaceous Yixian Formation (China) is published by Hethke et al. (2019).[34]
  • A study on the oxygen isotope composition of whale barnacle shells from three Pleistocene localities along the eastern Pacific coast, and on their implications for the knowledge of the history of whale migrations, is published by Taylor et al. (2019).[35]
  • A study on the internal soft-tissue anatomy of Hesslandona angustata is published by Liu et al. (2019).[36]

New taxa[edit]

Malacostracans[edit]

Name Novelty Status Authors Age Type locality Country Notes Images

Acanthocarpus osborni[37]

Sp. nov

Valid

Feldmann, Schweitzer & Phillips

Oligocene

 United States

A crab, a species of Acanthocarpus.

Ahazianassa[38]

Gen. et comb. nov

Valid

Karasawa et al.

Late Cretaceous (CampanianMaastrichtian)

Izumi Group

 Japan

A member of the family Gourretiidae; a new genus for "Callianassa" masanorii Karasawa (1998).

Alontecarcinus[39]

Gen. et sp. nov

Valid

De Angeli & Caporiondo

Eocene (Bartonian)

 Italy

A crab belonging to the family Potamidae. The type species is A. buratoi.

Amatukamius[40]

Gen. et comb. nov

Valid

Karasawa & Ohara

Early Cretaceous (Barremian)

Arida Formation

 Japan

A member of Axiidea belonging to the family Micheleidae; a new genus for "Callianassa" sakakuraorum Karasawa (2000).

Anapagurus dalessandroi[41] Sp. nov Valid Beschin, Busulini & Tessier in Beschin et al. Eocene (Priabonian)  Italy A member of the family Paguridae.

Annuntidiogenes elongatus[42]

Sp. nov

Valid

Fraaije et al.

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A hermit crab belonging to the family Annuntidiogenidae.

Annuntidiogenes hoelderi[42]

Sp. nov

Valid

Fraaije et al.

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A hermit crab belonging to the family Annuntidiogenidae.

Anoplocarcinus[43]

Gen. et sp. nov

Valid

Schweitzer et al.

Early Cretaceous (Albian)

Hudspeth Formation

 United States

A crab belonging to the group Raninoida and to the family Orithopsidae. The type species is A. hudspethi.

Aragolambrus[44]

Gen. et sp. nov

Valid

Ferratges, Zamora & Aurell

Eocene (Ypresian)

 Spain

A crab belonging to the family Parthenopidae. Genus includes new species A. collinsi.

Archaeochiapasa[45]

Gen. et sp. nov

Valid

Guinot, Carbot-Chanona & Vega

Late Cretaceous (Cenomanian)

Sierra Madre Formation

 Mexico

A crab belonging to the group Heterotremata, assigned to the new family Archaeochiapasidae. The type species is A. mardoqueoi.

Archaeopus burgessi[46]

Sp. nov

Valid

Nyborg et al.

Late Cretaceous (Campanian)

Cedar District Formation

 Canada

A crab belonging to the family Retroplumidae.

Archaeopus diabolus[47]

Sp. nov

Valid

Schweitzer et al.

Late Cretaceous

 United States

Archaeopus hoploserratus[46]

Sp. nov

Valid

Nyborg et al.

Late Cretaceous (Campanian)

Northumberland Formation

 Canada

A crab belonging to the family Retroplumidae.

Archaeopus morenoensis[46]

Sp. nov

Valid

Nyborg et al.

Late Cretaceous (Maastrichtian)

Moreno Formation

 United States

A crab belonging to the family Retroplumidae.

Bachmayerus gasparici[42]

Sp. nov

Valid

Fraaije et al.

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A hermit crab belonging to the family Diogenidae.

Bachmayerus matushyznyi[42]

Sp. nov

Valid

Fraaije et al.

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A hermit crab belonging to the family Diogenidae.

Bahiacaris[48]

Gen. et comb. nov

Valid

Schweitzer et al.

Cretaceous

 Brazil

A caridean shrimp; a new genus for "Atyoida" roxoi Beurlen (1950). Announced in 2013; validated in 2023.[49]

Bathycalliax otsuka[50]

Sp. nov

Valid

Ando, Kawano & Ugai

Eocene

Sakasegawa Formation

 Japan

A member of Axiidea belonging to the family Bathycalliacidae.

Bolcacalliax[51]

Gen. et comb. nov

Valid

Pasini et al.

Eocene (Ypresian)

 Italy

A member of the family Callianassidae; a new genus for "Protaxius" eocenicus Secrétan (1975).

Brachynotus multituberculatus[41] Sp. nov Valid Beschin, Busulini & Tessier in Beschin et al. Eocene (Priabonian)  Italy A species of Brachynotus.

Calappilia gableorum[37]

Sp. nov

Valid

Feldmann, Schweitzer & Phillips

Oligocene

 United States

A species of Calappilia.

Calappilia granulosa[37]

Sp. nov

Valid

Feldmann, Schweitzer & Phillips

Oligocene

 United States

A species of Calappilia.

Calappilia guttata[52]

Sp. nov

Valid

Pasini, Garassino & Damarco

Oligocene (Rupelian)

 Italy

A species of Calappilia.

Calappilia perreaulti[37]

Sp. nov

Valid

Feldmann, Schweitzer & Phillips

Oligocene

 United States

A species of Calappilia.

Callichimaera[53]

Gen. et sp. nov

Valid

Luque et al.

Late Cretaceous (Cenomanian to Turonian)

Churuvita Group
Frontier Formation

 Colombia
 United States

A crab belonging to the new family Callichimaeridae. The type species is C. perplexa.

Caloxanthus kavanaghi[54]

Sp. nov

Valid

Collins, Garvie & Mellish

Cretaceous (Albian/Cenomanian)

Del Rio Formation

 United States

A crab belonging to the group Etyoidea and to the family Feldmannidae.

Cenocorystes libbyae[55]

Sp. nov

Valid

Van Bakel et al.

Late Cretaceous (Maastrichtian)

Peedee Formation

 United States

A palaeocorystoid crab.

Corallomursia prominens[41] Sp. nov Valid Beschin, Busulini & Tessier in Beschin et al. Eocene (Priabonian)  Italy A member of the family Calappidae.

Costadromia[56]

Gen. et sp. nov

Valid

Feldmann & Schweitzer

Late Cretaceous (late Campanian)

Wenonah Formation

 United States

A sponge crab. The type species is C. hajzeri.

Cretainermis[57]

Gen. et sp. nov

Valid

Prado, Calado & Barreto

Early Cretaceous (Aptian)

Romualdo Formation

 Brazil

A member of Penaeoidea. Genus includes new species C. pernambucensis.

Cretamaja klompmakeri[58]

Sp. nov

Valid

González-León & Moreno-Bedmar in Vega, González-León & Moreno-Bedmar

Early Cretaceous (Barremian)

Zapotitlan Formation

 Mexico

A crab.

Cretatrizocheles doerflesensis[42]

Sp. nov

Valid

Fraaije et al.

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A hermit crab belonging to the family Pylochelidae.

Cymodoce (?) scopula[59]

Sp. nov

Valid

Fraaije et al.

Early Cretaceous (Albian)

Basque-Cantabrian Basin

 Spain

A sphaeromatid isopod.

Daragrapsus setalis[41] Sp. nov Valid Beschin, Busulini & Tessier in Beschin et al. Eocene (Priabonian)  Italy A member of Grapsoidea of uncertain phylogenetic placement.

Dardanus caporiondoi[60]

Sp. nov

Valid

Ceccon & De Angeli

Early Oligocene

 Italy

A species of Dardanus.

Devonoaxius[61]

Gen. et sp. nov

Valid

Feldmann & Schweitzer

Devonian (Famennian)

Ohio Shale

 United States

A member of Axiidea. Genus includes new species D. garlandi.

Dinocarcinus[62]

Gen. et sp. nov

Valid

Van Bakel, Hyžný, Valentin & Robin in Robin et al.

Late Cretaceous (late Campanian)

 France

A crab, a member of Portunoidea sensu lato. The type species is D. velauciensis. Announced in 2019; validated in 2023.[63]

Doraranina[64]

Gen. et sp. nov

Valid

Gustafson, Nyborg & van Bakel

Early Eocene

Lookingglass Formation

 United States

A crab belonging to the group Raninoidea and to the family Lyreididae. Genus includes new species D. manleyi.

Dromiopsis marginospinosa[41] Sp. nov Valid Beschin, Busulini & Tessier in Beschin et al. Eocene (Priabonian)  Italy A member of the family Dromiidae.

Dromiopsis oscari[65]

Sp. nov

Valid

Nyborg et al.

Early Cretaceous (Albian)

Hudspeth Formation

 United States

?Dromiopsis rocchettii[41] Sp. nov Valid Beschin, Busulini & Tessier in Beschin et al. Eocene (Priabonian)  Italy A member of the family Dromiidae.

Ebalia pumicosa[60]

Sp. nov

Valid

Ceccon & De Angeli

Early Oligocene

 Italy

A species of Ebalia.

Eobodotria[66]

Gen. et sp. nov

Valid

Luque & Gerken

Late Cretaceous (CenomanianTuronian)

 Colombia

A member of Cumacea. Genus includes new species E. muisca.

Eocarpilius vicetinus[60]

Sp. nov

Valid

Ceccon & De Angeli

Early Oligocene

 Italy

A crab belonging to the group Carpilioidea and the family Carpiliidae.

Eodorippe connori[65]

Sp. nov

Valid

Nyborg et al.

Early Cretaceous (Albian)

Hudspeth Formation

 United States

A crab belonging to the superfamily Dorippoidea and the family Telamonocarcinidae. Originally described as a species of Eodorippe, but subsequently transferred to the genus Personadorippe.[67]

Eopaguropsis grandis[42]

Sp. nov

Valid

Fraaije et al.

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A hermit crab belonging to the family Diogenidae.

Eopaguropsis schindewolfi[42]

Sp. nov

Valid

Fraaije et al.

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A hermit crab belonging to the family Diogenidae.

Eucalliax capsulasetaea[68]

Sp. nov

Valid

Kiel et al.

Early Oligocene

Talara Basin

 Peru

A member of the family Callianassidae.

Eumunida veronensis[41] Sp. nov Valid Beschin, Busulini & Tessier in Beschin et al. Eocene (Priabonian)  Italy A member of the family Munididae. Originally described as a species of Eumunida, but subsequently transferred to the genus Muellermunida.[69]
Folguerolesia collinsi[41] Sp. nov Valid Beschin, Busulini & Tessier in Beschin et al. Eocene (Priabonian)  Italy A member of Leucosioidea belonging to the family Folguerolesiidae.

?Galathea genesis[70]

Sp. nov

Valid

Robins & Klompmaker

Late Jurassic

Ernstbrunn Limestone

 Austria

A member of the family Galatheidae.

Galatheites britmelanarum[70]

Sp. nov

Valid

Robins & Klompmaker

Late Jurassic

Ernstbrunn Limestone

 Austria

A member of Galatheoidea belonging to the family Catillogalatheidae.

Gastrosacus raboeufi[71]

Sp. nov

Valid

Fraaije et al.

Middle Jurassic (Bathonian)

 France

A member of Galatheoidea.

Glaessnerella yamoori[65]

Sp. nov

Valid

Nyborg et al.

Early Cretaceous (Albian)

Hudspeth Formation

 United States

Glabropilumnus bizzarinii[41] Sp. nov Valid Beschin, Busulini & Tessier in Beschin et al. Eocene (Priabonian)  Italy A member of the family Pilumnidae.

Glabropilumnus tuberculatus[60]

Sp. nov

Valid

Ceccon & De Angeli

Early Oligocene

 Italy

A crab belonging to the group Pilumnoidea and the family Pilumnidae.

Gyroplanus[47]

Gen. et sp. nov

Valid

Schweitzer et al.

Late Cretaceous

 United States

A cyclodorippoid crab. Genus includes new species G. sapphirinus.

Hepatiscus meneghinii[41] Sp. nov Valid Beschin, Busulini & Tessier in Beschin et al. Eocene (Priabonian)  Italy A member of the family Aethridae.

Hinecaris[38]

Gen. et sp. nov

Valid

Karasawa et al.

Late Cretaceous (CampanianMaastrichtian)

Izumi Group

 Japan

A member of the family Axiidae. The type species is H. simplex.

Kromtitis lluisprietoi[72]

Sp. nov

Valid

Ossó

Eocene (Lutetian)

 Spain

A crab belonging to the family Dynomenidae.

Lindahomola[73]

Gen. et sp. nov

Valid

Garassino et al.

Late Cretaceous (Maastrichtian)

Peedee Formation

 United States

A crab belonging to the family Homolidae. Genus includes new species L. longispina.

Linuparus shacklefordae[47]

Sp. nov

Valid

Schweitzer et al.

Late Cretaceous

 United States

A palinurid lobster.

Lophoranina lincki[37]

Sp. nov

Valid

Feldmann, Schweitzer & Phillips

Oligocene

 United States

A crab.

Macroacaena hannya[50]

Sp. nov

Valid

Ando, Kawano & Ugai

Eocene

Sakasegawa Formation

 Japan

A crab belonging to the family Lyreididae.

Maja prealpina[60]

Sp. nov

Valid

Ceccon & De Angeli

Early Oligocene

 Italy

A species of Maja.

Matutites periosus[37]

Sp. nov

Valid

Feldmann, Schweitzer & Phillips

Eocene

 United States

A crab.

Mesoparapylocheles strouhali[42]

Sp. nov

Valid

Fraaije et al.

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A hermit crab belonging to the family Parapylochelidae.

Mesoparapylocheles zapfei[42]

Sp. nov

Valid

Fraaije et al.

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A hermit crab belonging to the family Parapylochelidae.

Mesostylus pervesleri[74]

Sp. nov

Valid

Hyžný & Summesberger

Late Cretaceous (Maastrichtian)

Piesting Formation

 Austria

A member of the family Callianassidae.

Montemagambrus[75]

Gen. et sp. nov

Valid

Ceccon & De Angeli

Early Eocene

 Italy

A crab belonging to the family Parthenopidae. Genus includes new species M. corallinus.

Munidopsis salinaria[76]

Sp. nov

Valid

Gašparič et al.

Miocene (Langhian)

Tuzla Basin

 Bosnia and Herzegovina

A species of Munidopsis.

Munitheites kuepperi[42]

Sp. nov

Valid

Fraaije et al.

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A hermit crab belonging to the family Schobertellidae.

Nachioides[41] Gen. et sp. nov Valid Beschin, Busulini & Tessier in Beschin et al. Eocene (Priabonian)  Italy A member of the family Inachoididae. The type species is N. tuberculatus.

Natatolana poblana[77]

Sp. nov

Valid

Vega & Bruce in Vega et al.

Early Cretaceous (Valanginian-Hauterivian)

San Juan Raya Formation

 Mexico

An isopod belonging to the family Cirolanidae.

Nicolisia[41] Gen. et sp. nov Valid Beschin, Busulini & Tessier in Beschin et al. Eocene (Priabonian)  Italy A member of Xanthoidea belonging to the family Eumedonidae. The type species is N. antiqua.

Notopus adinae[37]

Sp. nov

Valid

Feldmann, Schweitzer & Phillips

Oligocene

 United States

A crab, a species of Notopus.

Ovalopus[70]

Gen. et comb. nov

Valid

Robins & Klompmaker

Late Jurassic

Ernstbrunn Limestone

 Austria
 Czech Republic
 Romania

A crab belonging to the group Homolodromioidea and to the family Jurellanidae. The type species is O. hoheneggeri (Moericke, 1889); genus also includes Coelopus convexus Schweitzer & Feldmann (2010) and Coelopus repandus Schweitzer & Feldmann (2010).

Pachycheles bazzanii[41] Sp. nov Valid Beschin, Busulini & Tessier in Beschin et al. Eocene (Priabonian)  Italy A member of the family Porcellanidae.

Pagurus haasei[78]

Sp. nov

Valid

Polkowsky & Fraaije

Oligocene (Chattian)

 Germany

A hermit crab, a species of Pagurus.

Palaemon aestuarius[79]

Sp. nov

Du et al.

Early Miocene

Mexican amber

 Mexico

A species of Palaemon.

Palaeopalinurus serratus[59]

Sp. nov

Valid

Fraaije et al.

Early Cretaceous (Albian)

Basque-Cantabrian Basin

 Spain

A spiny lobster.

Palinurus? sauerweinorum[47]

Sp. nov

Valid

Schweitzer et al.

Late Cretaceous

 United States

A palinurid lobster.

Panopeus santurbanensis[60]

Sp. nov

Valid

Ceccon & De Angeli

Early Oligocene

 Italy

A species of Panopeus.

Paronachoides[41] Gen. et sp. nov Valid Beschin, Busulini & Tessier in Beschin et al. Eocene (Priabonian)  Italy A member of the family Inachoididae. The type species is P. scaber.

Percnon santurbanensis[80]

Sp. nov

Valid

Ceccon & de Angeli

Oligocene

 Italy

A crab, a species of Percnon.

Petrolisthes punctatus[41] Sp. nov Valid Beschin, Busulini & Tessier in Beschin et al. Eocene (Priabonian)  Italy A species of Petrolisthes.

Pilumnopeus priscus[60]

Sp. nov

Valid

Ceccon & De Angeli

Early Oligocene

 Italy

A crab belonging to the group Pilumnoidea and the family Pilumnidae.

Pilumnus pedemontanus[60]

Sp. nov

Valid

Ceccon & De Angeli

Early Oligocene

 Italy

A species of Pilumnus.

Planoprosopon ogawaense[81]

Sp. nov

Valid

Karasawa & Hirota

Late JurassicEarly Cretaceous

Yatsuji Formation

 Japan

A crab belonging to the group Homolodromioidea and to the family Longodromitidae.

Planoprosopon sarumaru[81]

Sp. nov

Valid

Karasawa & Hirota

Late JurassicEarly Cretaceous

Yatsuji Formation

 Japan

A crab belonging to the group Homolodromioidea and to the family Longodromitidae.

Pretrizocheles[42]

Gen. et sp. nov

Valid

Fraaije et al.

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A hermit crab belonging to the family Pylochelidae. The type species is P. cocullo.

Prolecythocaris rieberi[82]

Sp. nov

Valid

Schweigert

Late Jurassic

 Germany

A crab belonging to the group Dromiacea and to the family Lecythocaridae.

Protopagurus[42]

Gen. et sp. nov

Valid

Fraaije et al.

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A hermit crab belonging to the family Paguridae. The type species is P. janoscheki.

Pseudocarinocarcinoides[41] Gen. et sp. nov Valid Beschin, Busulini & Tessier in Beschin et al. Eocene (Priabonian)  Italy A member of Goneplacoidea belonging to the family Carinocarcinoididae. The type species is P. karasawai.

Retropluma minuta[76]

Sp. nov

Valid

Gašparič et al.

Miocene (Langhian)

Tuzla Basin

 Bosnia and Herzegovina

A species of Retropluma.

Ripleycarcinus[47]

Gen. et sp. nov

Valid

Schweitzer et al.

Late Cretaceous

 United States

A homoloid crab. Genus includes new species R. primaevus.

Ripleycorystes[47]

Gen. et sp. nov

Valid

Schweitzer et al.

Late Cretaceous

 United States

A palaeocorystoid crab. Genus includes new species R. cardwelli.

Symethoides danieli[47]

Sp. nov

Valid

Schweitzer et al.

Late Cretaceous

 United States

A raninoid crab.

Tethysmunida setifera[41] Sp. nov Valid Beschin, Busulini & Tessier in Beschin et al. Eocene (Priabonian)  Italy A member of the family Munidopsidae.

Tithopaguristes[42]

Gen. et sp. nov

Valid

Fraaije et al.

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A hermit crab belonging to the family Diogenidae. The type species is T. porosus.

Tropidicarcinus[47]

Gen. et 2 sp. nov

Valid

Schweitzer et al.

Late Cretaceous

 United States

A dakoticancroid crab. Genus includes new species T. mcanallyae and T. starnesi.

Ululapagurus oroszyi[42]

Sp. nov

Valid

Fraaije et al.

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A hermit crab belonging to the family Diogenidae.

Vacuotheca[83]

Gen. et sp. nov

Valid

Schädel, Perrichot & Haug

Late Cretaceous (Turonian)

 France

A member of Isopoda belonging to the group Epicaridea. The type species is V. dupeorum.

Valamunida[84]

Gen. et sp. nov

Valid

Klompmaker & Robins in Hryniewicz et al.

Late Paleocene

Basilika Formation

 Norway

A member of the family Munididae. The type species is V. haeggi.

Verarena[41] Gen. et sp. nov Valid Beschin, Busulini & Tessier in Beschin et al. Eocene (Priabonian)  Italy A member of the family Euryplacidae. The type species is V. katoi.

Vibrissalana[70]

Gen. et sp. nov

Valid

Robins & Klompmaker

Late Jurassic

Ernstbrunn Limestone

 Austria

A porcelain crab. Genus includes new species V. jurassica.

Vicluvia[85]

Gen. et sp. nov

Valid

Larghi et al.

Middle Triassic (Ladinian)

Meride Limestone

 Italy
  Switzerland

A member of Lophogastrida. Genus includes new species V. lombardoae.

Ostracods[edit]

Name Novelty Status Authors Age Type locality Country Notes Images

Amicytheridea bilthanaensis[86]

Sp. nov

Valid

Chaudhary & Nagori

Late Cretaceous

Bagh Formation

 India

Bairdiocypris fabiformis[87]

Sp. nov

Valid

Melnikova

Late Ordovician

Delingde Formation

 Russia

Bairdoppilata dorsoarcuata[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Late Oligocene to early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Bairdiidae.

Bairdoppilata pintoi[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Late Oligocene to early Miocene

La Rosa Formation
Pirabas Formation

 Brazil
 Venezuela

A member of Podocopida belonging to the family Bairdiidae.

Bairdoppilata turonica[88]

Sp. nov

Valid

Slipper

Late Cretaceous (Turonian)

 United Kingdom

Bairdoppilata vandenboldi[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Late Oligocene to early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Bairdiidae.

Bakunella anae[89]

Sp. nov

Valid

Spadi et al.

Late MiocenePliocene

 Croatia

A member of the family Candonidae.

Bassleratia torquata[87]

Sp. nov

Valid

Melnikova

Late Ordovician

Delingde Formation

 Russia

Bolbozoe psittaca[90]

Sp. nov

Valid

Perrier et al.

Silurian

 United Kingdom

A member of Myodocopa.

Boreobolbina morkokiana[87]

Sp. nov

Valid

Melnikova

Late Ordovician

Delingde Formation

 Russia

Bradleya majorani[91]

Sp. nov

Valid

Bergue, Brandão & Anjos Zerfass

Pleistocene

Atlantic Ocean (Rio Grande Rise)

A member of the family Thaerocytheridae.

Bradleya ybate[91]

Sp. nov

Valid

Bergue, Brandão & Anjos Zerfass

Late Miocene

Atlantic Ocean (Rio Grande Rise)

A member of the family Thaerocytheridae.

Bythocypris wangi[92]

Sp. nov

Valid

Tanaka, Siveter & Williams

Devonian

Fukuji Formation

 Japan

Bythoceratina? asulcata[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Bythocytheridae.

Bythoceratina? sinuocostata[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Late Oligocene to early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Bythocytheridae.

Cardobairdia longitecta[88]

Sp. nov

Valid

Slipper

Late Cretaceous (Turonian)

 United Kingdom

Carinobairdia cabralae[93]

Sp. nov

Forel in Forel, Thuy & Wisshak

Late Triassic (Carnian)

Maantang Formation

 China

A member of the family Bairdiidae.

Cativella primaveraensis[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Trachyleberididae.

Clavofabellina fukujiensis[92]

Sp. nov

Valid

Tanaka, Siveter & Williams

Devonian

Fukuji Formation

 Japan

Clintiella antifrigga[94]

Sp. nov

Valid

Siveter et al.

Silurian

Hitoegane Formation

 Japan

A beyrichioid palaeocopid ostracod.

Coelochilina reticulata[87]

Sp. nov

Valid

Melnikova

Late Ordovician

Delingde Formation

 Russia

Costa spinaventralis[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Trachyleberididae.

Curfsina coarctata[86]

Sp. nov

Valid

Chaudhary & Nagori

Late Cretaceous

Bagh Formation

 India

Curfsina hanumanpuraensis[86]

Sp. nov

Valid

Chaudhary & Nagori

Late Cretaceous

Bagh Formation

 India

Cushmanidea? bragantinaensis[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Late Oligocene to early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Cushmanideidae.

Cyprideis? carmoi[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Late Oligocene to early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Cytherideidae.

Cytherella altacaelateralis[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Late Oligocene to early Miocene

Pirabas Formation

 Brazil

A member of the family Cytherellidae.

Cytherella kempfi[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Late Oligocene to early Miocene

Pirabas Formation

 Brazil

A member of the family Cytherellidae.

Cytherella notossinuosa[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Late Oligocene to early Miocene

Pirabas Formation

 Brazil

A member of the family Cytherellidae.

Cytherella truncatoides[88]

Sp. nov

Valid

Slipper

Late Cretaceous (Turonian)

 United Kingdom

Cytherella vulna[88]

Sp. nov

Valid

Slipper

Late Cretaceous (Turonian)

 United Kingdom

Cytherella weaveri[88]

Sp. nov

Valid

Slipper

Late Cretaceous (Turonian)

 United Kingdom

Cytherelloidea colini[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Early Miocene

Pirabas Formation

 Brazil

A member of the family Cytherellidae.

Cytherelloidea cumpassorobusta[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Early Miocene

Pirabas Formation

 Brazil

A member of the family Cytherellidae.

Cytherelloidea granulosa parca[88]

Subsp. nov

Valid

Slipper

Late Cretaceous (Turonian)

 United Kingdom

Cytherelloidea mediocythara[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Early Miocene

Pirabas Formation

 Brazil

A member of the family Cytherellidae.

Cytherelloidea quasilenisa[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Early Miocene

Pirabas Formation

 Brazil

A member of the family Cytherellidae.

Cytheretta fortiscostata[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Cytherettidae.

Cytheretta petrii[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Cytherettidae.

Cytheropteron hanumanpuraensis[86]

Sp. nov

Valid

Chaudhary & Nagori

Late Cretaceous

Bagh Formation

 India

Cytheropteron ratitalaiensis[86]

Sp. nov

Valid

Chaudhary & Nagori

Late Cretaceous

Bagh Formation

 India

Cytherura? ornatareticulum[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Cytheruridae.

Cytherura? punctocentrata[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Cytheruridae.

Cytherura? quasilenisa[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Late Oligocene to early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Cytheruridae.

Eocytheropteron bilthanaensis[86]

Sp. nov

Valid

Chaudhary & Nagori

Late Cretaceous

Bagh Formation

 India

Gibbosocythere[95]

Gen. et 2 sp. nov

Valid

Karpuk, Whatley & Maybury

Early Cretaceous

Crimean Peninsula

A member of Paradoxostomatidae. Genus includes new species G. ferelevis and G. cellulata.

Hemicytherura bradyisimilis[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Cytheruridae.

Hiatobairdia senegasi[93]

Sp. nov

Forel in Forel, Thuy & Wisshak

Late Triassic (Carnian and Norian)

Kuahongdong Formation
Maantang Formation

 China

A member of the family Bairdiidae.

Hiatobairdia zhengshuyingi[93]

Sp. nov

Forel in Forel, Thuy & Wisshak

Late Triassic (Carnian and Norian)

Kuahongdong Formation
Maantang Formation

 China

A member of the family Bairdiidae.

Hollinella orienta[94]

Sp. nov

Valid

Siveter et al.

Silurian

Gionyama Formation

 Japan

A hollinoid palaeocopid ostracod.

Hulingsina? lorenesmithae[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Cushmanideidae.

Hulingsina reticulorugosa[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Late Oligocene to early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Cushmanideidae.

Hungarella gommerii[93]

Sp. nov

Forel in Forel, Thuy & Wisshak

Late Triassic (Carnian)

Maantang Formation

 China

A member of the family Healdiidae.

Jatella ampla[87]

Sp. nov

Valid

Melnikova

Late Ordovician

Delingde Formation

 Russia

Krithe araucoensis[96]

Sp. nov

Valid

Bergue, Coimbra & Finger

Early Miocene

 Chile

Krithe dawnpetersonae[96]

Sp. nov

Valid

Bergue, Coimbra & Finger

Early Miocene

 Chile

Krithe nerudai[96]

Sp. nov

Valid

Bergue, Coimbra & Finger

Early Miocene

 Chile

Leperditella paula[87]

Sp. nov

Valid

Melnikova

Late Ordovician

Delingde Formation

 Russia

Loxoconcha corpulenta[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Loxoconchidae.

Microcythere acuminata[91]

Sp. nov

Valid

Bergue, Brandão & Anjos Zerfass

Quaternary

Atlantic Ocean (Rio Grande Rise)

A member of the family Microcytheridae.

Moierina grata[87]

Sp. nov

Valid

Melnikova

Late Ordovician

Delingde Formation

 Russia

Mydionobairdia cauladelicata[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Bairdiidae.

Nigeroloxoconcha baghensis[86]

Sp. nov

Valid

Chaudhary & Nagori

Late Cretaceous

Bagh Formation

 India

Nigeroloxoconcha diluta[86]

Sp. nov

Valid

Chaudhary & Nagori

Late Cretaceous

Bagh Formation

 India

Nudator[90]

Gen. et 4 sp. nov

Valid

Perrier et al.

Silurian (Wenlock and Ludlow)

 Poland
 United Kingdom
 France?

A member of Myodocopa belonging to the family Entomozoidae. The type species is N. inflatus; genus also includes new species N. angiportatus, N. artumatus and N. elegantulus.[97]

Ochesaarina aculeata[87]

Sp. nov

Valid

Melnikova

Late Ordovician

Delingde Formation

 Russia

Oculparva[90]

Gen. et sp. nov

Valid

Perrier et al.

Silurian

 United Kingdom

A member of Myodocopa. Genus includes new species O. moncola.

Ovocytheridea baghensis[86]

Sp. nov

Valid

Chaudhary & Nagori

Late Cretaceous

Bagh Formation

 India

Paijenborchella jeerabadensis[86]

Sp. nov

Valid

Chaudhary & Nagori

Late Cretaceous

Bagh Formation

 India

Pauproles[94]

Gen. et sp. nov

Valid

Siveter et al.

Silurian

Hitoegane Formation

 Japan

A eurychilinoid palaeocopid ostracod. Genus includes new species P. supparata.

Piscarista[90]

Gen. et sp. nov

Valid

Perrier et al.

Silurian

 United Kingdom

A member of Myodocopa. Genus includes new species P. sagenata.

Planiglandites[87]

Gen. et sp. nov

Valid

Melnikova

Late Ordovician

Delingde Formation

 Russia

Genus includes new species P. mirabilis.

Pontocyprella goussardi[93]

Sp. nov

Forel in Forel, Thuy & Wisshak

Late Triassic (Carnian)

Maantang Formation

 China

A member of the family Pontocyprididae.

Pontocyprella robusta cometa[88]

Subsp. nov

Valid

Slipper

Late Cretaceous (Turonian)

 United Kingdom

Pontocypris aguilerai[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Late Oligocene to early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Pontocypridae.

Pseudotethysia[95]

Gen. et sp. nov

Valid

Karpuk, Whatley & Maybury

Early Cretaceous

Crimean Peninsula

A member of Cytheruridae. Genus includes new species P. reticulata.

Quadracythere adornata[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Hemicytheridae.

Quadracythere fortisrobusta[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Hemicytheridae.

Quadracythere limbimodesta[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Hemicytheridae.

Rectella cistelliformis[87]

Sp. nov

Valid

Melnikova

Late Ordovician

Delingde Formation

 Russia

Retiprimites alveolatus[87]

Sp. nov

Valid

Melnikova

Late Ordovician

Delingde Formation

 Russia

Retiprimites mirandus[87]

Sp. nov

Valid

Melnikova

Late Ordovician

Delingde Formation

 Russia

Rostrocytheridea decurtata[98]

Sp. nov

Valid

Chaudhary, Nagori & Khosla

Late Cretaceous

Bagh Formation

 India

A member of the family Cytherideidae.

Rostrocytheridea divergens[98]

Sp. nov

Valid

Chaudhary, Nagori & Khosla

Late Cretaceous

Bagh Formation

 India

A member of the family Cytherideidae.

Semicytherura? fortisreticulata[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Cytheruridae.

Seminova[90]

Gen. et comb. nov

Valid

Perrier et al.

Silurian

 United Kingdom

A member of Myodocopa belonging to the superfamily Bolbozoidea. The type species is "Entomis" depressa Jones (1884).

Tenedocythere? rugosocostata[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Hemicytheridae.

Touroconcha ishizakii[30]

Sp. nov

Valid

Nogueira, Ramos & Hunt

Early Miocene

Pirabas Formation

 Brazil

A member of Podocopida belonging to the family Loxoconchidae.

Uthoernia sibirica[87]

Sp. nov

Valid

Melnikova

Late Ordovician

Delingde Formation

 Russia

Other crustaceans[edit]

Name Novelty Status Authors Age Type locality Country Notes Images

Austromegabalanus carrioli[99]

Sp. nov

Valid

Collareta et al.

Early Miocene

Chilcatay Formation

 Peru

A barnacle.

Calmanostraca hassbergella[100]

Sp. nov

Wagner, Hauh & Haug

Late Triassic (Carnian)

Hassberge Formation

 Germany

A member of Branchiopoda belonging to the group Calmanostraca.

Cretiscalpellum matrioni[101]

Sp. nov

Valid

Gale

Early Cretaceous (Albian)

 France

A barnacle. Originally described as a species of Cretiscalpellum; Gale (2020) transferred it to the genus Witherscalpellum.[102]

Cretiscalpellum mutterlosei[101]

Sp. nov

Valid

Gale

Early Cretaceous (Hauterivian)

 Germany

A barnacle.

Halicyne oosterinkorum[103]

Sp. nov

Valid

Schweitzer, Feldmann & Schinker

Middle Triassic (Anisian)

 Netherlands

A member of Cyclida.

Jaegerscalpellum[101]

Gen. et sp. et comb. nov

Valid

Gale

Early Cretaceous (Hauterivian to Albian)

 Germany

A barnacle. Genus includes new species J. elegans, as well as J. comptum (Withers, 1910) and J. politum (Darwin, 1851).

Liapora[104]

Gen. et sp. nov

Valid

Keupp

Early Jurassic (Pliensbachian)

 Germany

Originally described as a scyphozoan polyp; genus includes new species L. neubigi. Gale, Keupp & Schweigert (2022) reinterpreted it as capitular valves of a phosphatic barnacle that had been described as Pollicipes(?) lotharingica by Méchin in 1901.[105]

Martillepas auriculum[101]

Sp. nov

Valid

Gale

Early Cretaceous (Hauterivian)

 Germany

A barnacle.

Martillepas decoratus[101]

Sp. nov

Valid

Gale

Early Cretaceous (Hauterivian)

 Germany

A barnacle.

Notostraca oleseni[100]

Sp. nov

Wagner, Hauh & Haug

Early Cretaceous

Yixian Formation

 China

A member of Branchiopoda belonging to the group Notostraca.

Paraleptestheria mitchelli[106]

Sp. nov

Valid

Gallego et al.

Late Eocene

 United Kingdom

A clam shrimp.

Pedupycnolepas pulcher[101]

Sp. nov

Valid

Gale

Early Cretaceous (Hauterivian)

 Germany

A barnacle.

Pemphilimnadiopsis cheni[107]

Sp. nov

Valid

Liao et al.

Carboniferous (Pennsylvanian)

Benxi Formation

 China

A clam shrimp.

Perumegabalanus[108]

Gen. et sp. nov

Valid

Coletti et al.

Miocene (Burdigalian)

Chilcatay Formation

 Peru

A barnacle. Genus includes new species P. calziai.

Platylepas mediterranea[109]

Sp. nov

Valid

Collareta et al.

Early Pleistocene (Gelasian)

 Italy

A barnacle, a species of Platylepas.

Prolatcyclus[110]

Gen. et comb. et sp. nov

Valid

Mychko et al.

Carboniferous (Viséan)

 Russia
 United Kingdom

A member of Cyclida. Genus includes P. martinensis (Goldring, 1967) and a new species P. kindzadza.

Verruca alabamensis[111]

Sp. nov

Valid

Perreault & Buckeridge

Paleogene

Yazoo Formation

 United States

A barnacle belonging to the family Verrucidae.

Verruca gailgoedertae[111]

Sp. nov

Valid

Perreault & Buckeridge

Middle Eocene

 United States

A barnacle belonging to the family Verrucidae.

Verruca sorrellae[111]

Sp. nov

Valid

Perreault & Buckeridge

Late Eocene to earliest Oligocene

 United States

A barnacle belonging to the family Verrucidae.

Insects[edit]

Trilobites[edit]

Research[edit]

  • A study on the rates of evolution of trilobites in the Cambrian is published by Paterson, Edgecombe & Lee (2019), who interpret their findings as indicative of Cambrian rather than Precambrian origin of trilobites, and as indicating that the Cambrian explosion had already largely concluded by the time the typical Cambrian fossil record begins (≈521 Ma).[112]
  • A study on repaired injuries in trilobite specimens from the Cambrian Ruin Wash Lagerstätte (Nevada, United States) is published by Pates & Bicknell (2019), who interpret their findings as indicative of species specific predation on Cambrian trilobites, and of enlarged spines in some trilobite species serving as possible predation deterrents.[113]
  • A study on the morphology and ontogeny of Changaspis elongata, based on data from specimens from the Lazizhai section of the Balang Formation (Cambrian Stage 4; Guizhou, China), is published by Du et al. (2019).[114]
  • A cluster of over a 100 juvenile trilobites belonging to the genus Aphelaspis is reported from the Cambrian (Paibian) Conasauga Formation (Georgia, United States) by Schwimmer & Montante (2019).[115]
  • A study on patterns of occupancy and diversity trajectories in late Cambrian-early Ordovician trilobite communities from the Argentine Cordillera Oriental is published by Serra, Balseiro & Waisfeld (2019).[116]
  • A study on the phylogenetic relationships of members of the family Olenidae is published by Monti & Confalonieri (2019).[117]
  • A study on the phylogenetic relationships of members of the olenid subfamily Balnibarbiinae is published by Hopkins (2019).[118]
  • Nine specimens of Symphysurus ebbestadi preserving all key stages of the exoskeleton moulting process are described from the Lower Ordovician Fezouata Formation (Morocco) by Drage et al. (2019).[119]
  • A study on clusters of Ampyx priscus from the Ordovician Fezouata Shale of Morocco is published by Vannier et al. (2019), who interpret these clusters as evidence of a collective behaviour rather than a result of passive transportation and accumulation.[120]
  • A study on the phylogenetic relationships of the species and subspecies assigned to the phacopid genus Austerops is published by Oudot et al. (2019).[121]
  • A study on the morphological changes in the ontogeny of the Ordovician phacopid trilobite Calyptaulax strasburgensis is published by Jacobs & Carlucci (2019).[122]
  • A well-preserved body cluster of 18 specimens of Arctinurus boltoni is described from the Silurian Rochester Shale (New York, United States) by Bicknell, Paterson & Hopkins (2019).[123]
  • A study on development and evolution of the cephalon and the pygidium in phacopid trilobites is published by Oudot et al. (2019).[124]
  • A study on the internal structures of eyes of trilobites belonging to the genera Asaphus and Archegonus is published by Scholtz, Staude & Dunlop (2019);[125] the study is subsequently criticized by Schoenemann & Clarkson (2021).[126][127]
  • A study on the variability of trilobite moulting behaviour is published by Drage (2019).[128]

New taxa[edit]

Name Novelty Status Authors Age Type locality Country Notes Images

Anacaenaspis yanpingensis[129]

Sp. nov

Valid

Wei & Zhan

Silurian (Rhuddanian)

Niuchang Formation

 China

Asaphellus intermedius[130]

Sp. nov

Valid

Ghobadi Pour

Ordovician

 Iran

Asteropyge boeckae[131]

Sp. nov

Valid

Van Viersen, Taghon & Magrean

Devonian (Eifelian)

Jemelle Formation

 Belgium

Asteropyge eonia[131]

Sp. nov

Valid

Van Viersen, Taghon & Magrean

Devonian (Eifelian)

Jemelle Formation

 France

Asteropyge filoxenia[131]

Sp. nov

Valid

Van Viersen, Taghon & Magrean

Devonian (Eifelian)

Jemelle Formation

 Belgium

Astycoryphe rugocauda[131]

Sp. nov

Valid

Van Viersen, Taghon & Magrean

Devonian (Eifelian)

Couvin Formation

 Belgium

A member of Proetida.

Astycoryphe tureki[132]

Sp. nov

Valid

Mergl & Budil

Devonian (Eifelian)

 Czech Republic

A member of Proetida.

Bigotina kelleri[133]

Sp. nov

Valid

Geyer

Cambrian Stage 3

Igoudine Formation

 Morocco

A member of Redlichiina belonging to the family Bigotinidae.

Bigotina monningeri[133]

Sp. nov

Valid

Geyer

Cambrian

 Morocco

A member of Redlichiina belonging to the family Bigotinidae.

Bigotinops chouberti[133]

Sp. nov

Valid

Geyer

Cambrian

 Morocco

A member of Redlichiina belonging to the family Bigotinidae.

Brachymetopus (Conimetopus) alekseevi[134]

Sp. nov

Valid

Mychko in Mychko & Savchuk

Early Permian

 Russia

Brevibole minerva[135]

Sp. nov

Valid

Müller & Hahn

Carboniferous (Mississippian)

 Germany

A member of the family Phillipsiidae belonging to the subfamily Achegoninae.

Bronteopsis tenuirhachis[136]

Sp. nov

Valid

Ove, Ebbestad & Fortey

Late Ordovician

Povorotnaya Formation

 Russia

A member of the family Styginidae.

Calymene dastanpouri[137]

Sp. nov

Valid

Ameri, Arjmandzadeh & Ghoorcgi

Early Silurian

Niur Formation

 Iran

Announced in 2019; the final version of the article naming was published in 2021.

Carbonoproetus bellona[135]

Sp. nov

Valid

Müller & Hahn

Carboniferous (Mississippian)

 Germany

A member of the family Phillipsiidae belonging to the subfamily Bollandiinae.

Carbonoproetus volturnus[135]

Sp. nov

Valid

Müller & Hahn

Carboniferous (Mississippian)

 Germany

A member of the family Phillipsiidae belonging to the subfamily Bollandiinae.

Conophrys multituberculatus[130]

Sp. nov

Valid

Ghobadi Pour

Ordovician

 Iran

Cyphaspides ammari[138]

Sp. nov

Valid

Chatterton, Gibb & McKellar

Devonian (Eifelian)

Bou Tchrafine Group

 Morocco

A member of Proetida belonging to the family Aulacopleuridae.

Cyphaspides nicoleae[138]

Sp. nov

Valid

Chatterton, Gibb & McKellar

Devonian (Eifelian)

Bou Tchrafine Group

 Morocco

A member of Proetida belonging to the family Aulacopleuridae.

Cyphaspides pankowskiorum[138]

Sp. nov

Valid

Chatterton, Gibb & McKellar

Devonian (Eifelian)

Bou Tchrafine Group

 Morocco

A member of Proetida belonging to the family Aulacopleuridae.

Cyphaspis insolata[131]

Sp. nov

Valid

Van Viersen, Taghon & Magrean

Devonian (Eifelian)

Jemelle Formation

 Belgium

Cyphaspis iuxta[131]

Sp. nov

Valid

Van Viersen, Taghon & Magrean

Devonian (Eifelian)

Jemelle Formation

 Belgium

Cyphaspis konstantini[139]

Sp. nov

Valid

Alberti

Devonian (Emsian)

 Germany

A member of the family Aulacopleuridae.

Cyrtoproetus korni[135]

Sp. nov

Valid

Müller & Hahn

Carboniferous (Mississippian)

 Germany

A member of the family Phillipsiidae belonging to the subfamily Achegoninae.

Cyrtoproetus roesenbeckensis[135]

Sp. nov

Valid

Müller & Hahn

Carboniferous (Mississippian)

 Germany

A member of the family Phillipsiidae belonging to the subfamily Achegoninae.

Cyrtosymbolinella[140]

Gen. et sp. nov

Valid

Basse & Lemke

Devonian

 Germany

A member of the family Phillipsiidae belonging to the subfamily Cyrtosymbolinae. The type species is C. svojtkai.

Debrenella[133]

Gen. et sp. nov

Valid

Geyer

Cambrian

 Morocco

A member of Redlichiida belonging to the family Fallotaspididae. Genus includes new species D. larvalis.

Diademaproetus pertinax[131]

Sp. nov

Valid

Van Viersen, Taghon & Magrean

Devonian (Eifelian)

Jemelle Formation

 Belgium

A member of Proetida.

Dionide trigintasegmentata[136]

Sp. nov

Valid

Ove, Ebbestad & Fortey

Late Ordovician

Barkov Formation

 Russia

A member of Asaphida belonging to the family Dionididae.

Eladiolinania[133]

Gen. et 2 sp. et comb. nov

Valid

Geyer

Cambrian

Igoudine Formation

 Morocco
 Spain?

A member of Redlichiina belonging to the family Bigotinidae. Genus includes new species E. castor and E. pollux, and possibly also "Serrania" palaciosi Liñán et al. (2008) and "Serrania" gordaensis Liñán et al. (2008).

Evenkaspis (Bilobaspis) mirabilis[136]

Subgen. et sp. nov

Valid

Ove, Ebbestad & Fortey

Late Ordovician

Tolmachev Formation

 Russia

A member of the family Pterygometopidae.

Failleana superba[136]

Sp. nov

Valid

Ove, Ebbestad & Fortey

Late Ordovician

Povorotnaya Formation

 Russia

A member of the family Styginidae.

Fallotaspis antecedens[133]

Sp. nov

Valid

Geyer

Cambrian

Igoudine Formation

 Morocco

A member of Redlichiida belonging to the family Fallotaspididae.

Geesops icovellaunae[131]

Sp. nov

Valid

Van Viersen, Taghon & Magrean

Devonian (Eifelian)

Jemelle Formation

 Belgium

Gerastos kesselaeri[131]

Sp. nov

Valid

Van Viersen, Taghon & Magrean

Devonian (Eifelian)

Jemelle Formation

 Belgium

Gerastos silvicultrix[131]

Sp. nov

Valid

Van Viersen, Taghon & Magrean

Devonian (Emsian)

Heisdorf Formation

 Germany

Heliopeltis morrisoni[141]

Sp. nov

Valid

Basse, Müller & Habenicht

Devonian (Emsian)

 Germany

A member of the family Scutelluidae belonging to the subfamily Kolihapeltinae.

Hollandiella[142]

Gen. et 3 sp. nov

Valid

Van Viersen & Lerouge

Devonian (Pragian and Emsian)

Ihandar Formation
Timrhanrhart Formation

 Morocco

A member of the family Proetidae belonging to the subfamily Cornuproetinae. The type species is H. curvirostris; genus also includes new species H. lebruni and H. verecunda.

Interproetus mizobuchii[143]

Sp. nov

Valid

Stocker et al.

Paleozoic

 Japan

A member of Proetida belonging to the family Proetidae.

Issendalenia[133]

Gen. et sp. nov

Valid

Geyer

Cambrian

Igoudine Formation

 Morocco

A member of Redlichiina belonging to the family Bigotinidae. Genus includes new species I. grandispina.

Kasachstania alperovichi[144]

Sp. nov

Valid

Ghobadi Pour et al.

Devonian (Lochkovian)

Shakshagaily Formation

 Kazakhstan

Kellerwaldops[140]

Gen. et sp. nov

Valid

Basse & Lemke

Devonian

 Germany

A member of the family Phacopidae belonging to the subfamily Phacopinae. The type species is K. kaufmanni.

Kettneraspis eftychia[131]

Sp. nov

Valid

Van Viersen, Taghon & Magrean

Devonian (Eifelian)

Jemelle Formation

 France

A member of the family Odontopleuridae.

Koneprusia chimaera[139]

Sp. nov

Valid

Alberti

Devonian (Emsian)

 Germany

A member of the family Odontopleuridae.

Koneprusites oresibios[142]

Sp. nov

Valid

Van Viersen & Lerouge

Devonian (Eifelian)

 Morocco

A member of the family Proetidae belonging to the subfamily Cornuproetinae.

Lepidoproetus arenicolus[142]

Sp. nov

Valid

Van Viersen & Lerouge

Devonian (Pragian)

 Morocco

A member of the family Proetidae belonging to the subfamily Cornuproetinae.

Liobole (Liobole) aurora[135]

Sp. nov

Valid

Müller & Hahn

Carboniferous (Mississippian)

 Germany

A member of the family Phillipsiidae belonging to the subfamily Achegoninae.

Liobole (Panibole) neptun[135]

Sp. nov

Valid

Müller & Hahn

Carboniferous (Mississippian)

 Germany

A member of the family Phillipsiidae belonging to the subfamily Achegoninae.

Nephranops (Maternia)[145]

Subgen. et sp. nov

Valid

Feist

Devonian (Famennian)

 France
 Germany

A member of Phacopida. The type species of the subgenus is "Nephranops incisus" dillanus Richter & Richter (1926) (raised to the rank of a separate species Nephranops (Maternia) dillanus); the subgenus also includes Nephranops franconicus Alberti (1970) and a new species Nephranops (Maternia) occitanicus.

Nephranops (Nephranops) angerae[145]

Sp. nov

Valid

Feist

Devonian (Famennian)

 France

A member of Phacopida.

Niobe (Niobella) obscura[146]

Sp. nov

Valid

Zhou & Zhou

Ordovician

Dawan Formation

 China

A member of the family Asaphidae belonging to the subfamily Isotelinae. Originally described as a species of Niobe, but subsequently transferred to the genus Zhiyia.[147]

Paralejurus colossicus[148]

Sp. nov

Valid

Alberti

Devonian (Emsian)

 Germany

A member of the family Scutelluidae.

Paralejurus flickorum[148]

Sp. nov

Valid

Alberti

Devonian (Emsian)

 Germany

A member of the family Scutelluidae.

Pararemopleurides ornatissimus[136]

Sp. nov

Valid

Ove, Ebbestad & Fortey

Late Ordovician

Povorotnaya Formation

 Russia

A member of Asaphida belonging to the family Remopleurididae.

Pedinopariops ceuthonymus[131]

Sp. nov

Valid

Van Viersen, Taghon & Magrean

Devonian (Eifelian)

Jemelle Formation

 France

A member of the family Phacopidae.

Pedinopariops richterianus geminus[131]

Subsp. nov

Valid

Van Viersen, Taghon & Magrean

Devonian (Eifelian)

Jemelle Formation

 Belgium

A member of the family Phacopidae.

Placoparia africana[149]

Sp. nov

In press

Pereira & Gutiérrez-Marco in Gutiérrez-Marco et al.

Ordovician (Sandbian)

 Morocco

Announced in 2019; the final version of the article naming it is not published yet.

Polypleuraspidella[150]

Gen. et comb. nov

Valid

Peel

Cambrian

Spence Shale

 United States
( Utah)

A new genus for "Polypleuraspis" cooperi Robison & Babcock (2011).

Polypleuraspis glacialis[150]

Sp. nov

Valid

Peel

Cambrian (Miaolingian)

Telt Bugt Formation

 Greenland

Pseudobigotina[133]

Gen. et sp. nov

Valid

Geyer

Cambrian

 Morocco

A member of Redlichiina belonging to the family Bigotinidae. Genus includes new species P. antiatlasensis.

Pulvinocephalus[145]

Gen. et comb. et 2 sp. et subsp. nov

Valid

Feist

Devonian (Famennian)

 Germany
 Russia

A member of Phacopida. The type species is "Phacops (Trimerocephalus)" steinachensis Richter & Richter (1926) (including a new subspecies Pulvinocephalus steinachensis triangulus); genus also includes "Phacops (Cryphops)" latilimbatus Maximova (1955) and new species Pulvinocephalus ovatus and Pulvinocephalus walliseri.

Radiaspis taghonorum[151]

Sp. nov

Valid

Van Viersen & Magrean

Middle Devonian

Hanonet Formation

 Belgium

A member of the family Odontopleuridae.

Raymondella plastron[136]

Sp. nov

Valid

Ove, Ebbestad & Fortey

Late Ordovician

Povorotnaya Formation

 Russia

A member of the family Raphiophoridae.

Redlichia rex[152]

Sp. nov

Valid

Holmes, Paterson & García-Bellido

Cambrian Stage 4

Emu Bay Shale

 Australia

Robergia nikolaiseni[136]

Sp. nov

Valid

Ove, Ebbestad & Fortey

Late Ordovician

Barkov Formation

 Russia

A member of Asaphida belonging to the family Remopleurididae.

Robergia subtilis[136]

Sp. nov

Valid

Ove, Ebbestad & Fortey

Late Ordovician

Povorotnaya Formation

 Russia

A member of Asaphida belonging to the family Remopleurididae.

Scabriscutellum archinalae[131]

Sp. nov

Valid

Van Viersen, Taghon & Magrean

Devonian (Eifelian)

Jemelle Formation

 Belgium

A member of the family Scutelluidae.

Sculptoproetus raki[142]

Sp. nov

Valid

Van Viersen & Lerouge

Devonian (Emsian)

 Morocco

A member of the family Proetidae belonging to the subfamily Cornuproetinae.

Septimopeltis akatastasia[131]

Sp. nov

Valid

Van Viersen, Taghon & Magrean

Devonian (Eifelian)

Jemelle Formation

 Belgium

A member of the family Scutelluidae.

Struveops wiedaensis[145]

Sp. nov

Valid

Feist

Devonian (Famennian)

 Germany

A member of Phacopida.

Suvorovaella[133]

Gen. et comb. nov

Valid

Geyer

Cambrian

El'gyan Formation

 Russia

A member of Redlichiina belonging to the family Bigotinidae. The type species is "Bigotinops" privus Suvorova (1960); genus might also include "Bigotinops" patrius Suvorova (1960).

Symphysurus ebbestadi[153]

Sp. nov

Valid

Gutierrez-Marco, Rabano & Garcia-Bellido

Ordovician (Tremadocian)

 Morocco

A member of the family Nileidae.

Tafilaltaspis rufa[142]

Sp. nov

Valid

Van Viersen & Lerouge

Devonian (Eifelian)

 Morocco

A member of the family Proetidae belonging to the subfamily Eremiproetinae.

Tesselacauda kriegerae[154]

Sp. nov

Valid

Adrain & Karim

Early Ordovician

Fillmore Formation
Garden City Formation

 United States

Tesselacauda morrisoni[154]

Sp. nov

Valid

Adrain & Karim

Early Ordovician

Fillmore Formation

 United States

Timsaloproetus gibbae[142]

Sp. nov

Valid

Van Viersen & Lerouge

Devonian (Emsian)

Khebchia Formation

 Morocco

A member of the family Proetidae belonging to the subfamily Cornuproetinae.

Timsaloproetus pulchistriatus[142]

Sp. nov

Valid

Van Viersen & Lerouge

Devonian (Emsian)

Timrhanrhart Formation

 Morocco

A member of the family Proetidae belonging to the subfamily Cornuproetinae.

Timsaloproetus weddigei[142]

Sp. nov

Valid

Van Viersen & Lerouge

Devonian (Eifelian)

Bicken-Ense Formation

 Germany

A member of the family Proetidae belonging to the subfamily Cornuproetinae.

Tioutella[133]

Gen. et sp. nov

Valid

Geyer

Cambrian

Igoudine Formation

 Morocco

A member of Redlichiina belonging to the family Bigotinidae. Genus includes new species T. floccofavosa.

Trifoliops septimanicus[145]

Sp. nov

Valid

Feist

Devonian (Famennian)

 France

A member of Phacopida.

Tropidocoryphe insciens[131]

Sp. nov

Valid

Van Viersen, Taghon & Magrean

Devonian (Eifelian)

Jemelle Formation

 Belgium

A member of Proetida.

Other arthropods[edit]

Research[edit]

  • A study on the morphology of appendages of the euarthropod Ercaicunia multinodosa from the Early Cambrian Chengjiang biota (China) is published by Zhai et al. (2019).[155]
  • Description of the appendicular morphology of Sinoburius lunaris is published by Chen et al. (2019).[156]
  • Description of a new specimen of Emeraldella brutoni from the Cambrian Wheeler Formation (Drum Mountains, Utah, United States), providing new information on the appendicular anatomy of this species, is published by Lerosey-Aubril & Ortega-Hernández (2019).[157]
  • Description of new bradoriid fossils from the Chengjiang biota, revealing hitherto unknown morphological differences in limb anatomy between bradoriid species, is published by Zhai et al. (2019), who evaluate the implications of this finding for the knowledge of the phylogenetic relationships of bradoriids.[158]
  • A study on the appendage structure of Naraoia spinosa at both juvenile and adult stages is published by Zhai et al. (2019).[159]
  • Description of exceptionally preserved soft tissues from mature individuals of Peronopsis and Ptychagnostus from the Cambrian Burgess Shale (British Columbia, Canada), and a study on their implications for inferring the phylogenetic placement of agnostinids within Arthropoda, is published by Moysiuk & Caron (2019).[160]
  • Completely enrolled, phosphatized specimens of Tsunyidiscus yanjiazhaiensis are described from Cambrian Stage 3 of South China by Dai et al. (2019).[161]
  • The ontogenetic series of Pagetia vinusta is documented from the Cambrian (Wuliuan) Kaili Formation (Guizhou, China) by Cui et al. (2019).[162]
  • A study reevaluating crystals of a copper sulfide mineral found in association with specimens of Marrella splendens from the Burgess Shale, originally interpreted as evidence that the animal's blood contained the Cu-bearing protein hemocyanin, is published by Gaines et al. (2019).[163]
  • A study on the anatomy of the nervous system of Alalcomenaeus is published by Ortega-Hernández, Lerosey-Aubril & Pates (2019).[164]
  • Juvenile specimens of stylonurine eurypterids belonging to the family Hardieopteridae are described from the Famennian locality of Strud (Belgium) by Lamsdell et al. (2019), who interpret their findings as evidence indicating that eurypterids underwent a marine to freshwater transition during the Devonian, and that juvenile eurypterids inhabited sheltered nursery pools (and migrated to rivers upon reaching maturity).[165]
  • A study on the structure of the eyes of eurypterids is published by Schoenemann, Poschmann & Clarkson (2019).[166]
  • A study on ontogenetic changes in prosomal morphology in Carboniferous horseshoe crab Euproops danae is published by Tashman, Feldmann & Schweitzer (2019).[167]
  • A study on the anatomy of the appendages of the Cretaceous xiphosurid Tachypleus syriacus and on the evolution of fossil horseshoe crab appendages is published by Bicknell et al. (2019).[168]
  • A study re-evaluating the fossil evidence for lateral compound eyes in Paleozoic horseshoe crabs is published by Bicknell, Amati & Ortega-Hernández (2019).[169]
  • A study on the morphology of the millipede Phryssonotus burmiticus from the Cretaceous amber from Myanmar is published by Su, Cai & Huang (2019).[170]

New taxa[edit]

Name Novelty Status Authors Age Type locality Country Notes Images

Albalimulus[171]

Gen. et sp. nov

Bicknell & Pates

Carboniferous (Tournaisian)

Ballagan Formation

 United Kingdom

A member of Limuloidea. The type species is A. bottoni.

Andrognathus burmiticus[172]

Sp. nov

Valid

Moritz & Wesener

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A millipede, a species of Andrognathus.

Ankitokazocaris tatensis[173]

Sp. nov

Disputed

Ehiro et al.

Early Triassic (Olenekian)

Osawa Formation

 Japan

A thylacocephalan. Laville et al. (2021) considered this species to be a junior synonym of Ankitokazocaris chaohuensis.[174]

Beckwithia? gracilis[175]

Sp. nov

Valid

Zhu, Lerosey-Aubril & Ortega-Hernández

Cambrian (Furongian)

Sandu Formation

 China

A member of Aglaspidida.

Burmanopetalum[176]

Gen. et sp. nov

Valid

Stoev, Moritz & Wesener

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A millipede belonging to the group Callipodida and to the new family Burmanopetalidae. The type species is B. inexpectatum.

Cambroraster falcatus[177]

Gen. et sp. nov

Valid

Moysiuk & Caron

Cambrian (Wuliuan)

Burgess Shale

 Canada

A member of Radiodonta belonging to the family Hurdiidae.

Concavicaris parva[173]

Sp. nov

Valid

Ehiro et al.

Early Triassic (Olenekian)

Osawa Formation

 Japan

A thylacocephalan. Originally described as a species of Concavicaris; Laville et al. (2021) transferred this species to the genus Ankitokazocaris.[174]

Duraznovis[178]

Gen. et sp. nov

Valid

Lara et al.

Late Triassic (Carnian)

Potrerillos Formation

 Argentina

An arthropod of uncertain phylogenetic placement. Genus includes new species D. gallegoi. It was argued to be an insect belonging to the group Cicadomorpha by Fu & Huang (2020).[179]

Electrosminthuridia[180]

Gen. et sp. nov

Robin, D'Haese & Barden

Miocene

Dominican amber

 Dominican Republic

A springtail belonging to the family Sminthurididae. The type species is E. helibionta.

Falcatacaris[181]

Gen. et sp. nov

Valid

Braig et al.

Late Jurassic (Tithonian)

Altmühltal Formation

 Germany

A thylacocephalan. The type species is F. bastelbergeri.

Fibulacaris[182]

Gen. et sp. nov

Valid

Izquierdo-López & Caron

Cambrian (Wuliuan)

Burgess Shale

 Canada

A bivalved arthropod belonging to the group Mandibulata. The type species is F. nereidis.

Hoplitaspis[183]

Gen. et sp. nov

Valid

Lamsdell, Gunderson & Meyer

Late Ordovician

Big Hill Formation

 United States

A member of Chasmataspidida belonging to the family Diploaspididae. The type species is H. hiawathai.

Hyleoglomeris groehni[184]

Sp. nov

Valid

Wesener

Eocene

Baltic amber

Europe (Baltic Sea region)

A millipede belonging to the family Glomeridae.

Ligulacaris[185]

Gen. et sp. nov

Valid

Charbonnier et al.

Early Triassic

 United States

A thylacocephalan. Genus includes new species L. parisiana.

Luohuilinella deletres[186]

Sp. nov

Valid

Hou et al.

Cambrian Stage 3

Chiungchussu Formation

 China

A probable member of Artiopoda and the order Xandarellida.

L. deletres (left)

Micragnostus iturbensis[187]

Sp. nov

Valid

Tortello & Esteban

Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

A member of Agnostida. Announced in 2019; the final version of the article naming it was published in 2020.

Miyagicaris[173]

Gen. et sp. nov

Disputed

Ehiro et al.

Early Triassic (Olenekian)

Osawa Formation

 Japan

A thylacocephalan. Genus includes new species M. costata. However, Laville et al. (2021) considered M. costata to be a junior synonym of Ankitokazocaris utatsuensis.[174]

Mollisonia plenovenatrix[188]

Sp. nov

Valid

Aria & Caron

Cambrian (Wuliuan)

Burgess Shale

 Canada

A member of Chelicerata belonging to the family Mollisoniidae.

Mongolarachne chaoyangensis[189]

Sp. nov

Disputed

Cheng et al.

Early Cretaceous

Yixian Formation

 China

An arthropod of uncertain phylogenetic placement. Originally described as a spider; Selden et al. (2019) reinterpreted it as a faked fossil based on a fossil crayfish, and considered it to be a probable junior synonym of Cricoidoscelosus aethus.[190]

Parisicaris[185]

Gen. et sp. nov

Disputed

Charbonnier et al.

Early Triassic

 United States

A thylacocephalan. Genus includes new species P. triassica. However, Laville et al. (2021) transferred P. triassica to the genus Ankitokazocaris.[174]

Pickettia[191]

Gen. et comb. nov

Valid

Bicknell, Lustri & Brougham

Late Devonian

 United States

A member of Xiphosura; a new genus for "Bellinurus" carteri Eller, 1940.

Pseudobeyrichona monile[192]

Sp. nov

Valid

Streng & Geyer

Cambrian

Tannenknock Formation

 Germany

A member of Bradoriida belonging to the family Hipponicharionidae.

Siphonophora hui[193]

Sp. nov

Valid

Jiang et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A millipede belonging to the group Siphonophorida.

Sloveniolimulus[194]

Gen. et sp. nov

Valid

Bicknell et al.

Middle Triassic (Anisian)

Strelovec Formation

 Slovenia

A horseshoe crab. Genus includes new species S. rudkini.

Stilpnocephalus[195]

Gen. et sp. nov

Valid

Selden, Simonetto & Marsiglio

Carboniferous (Kasimovian)

Meledis Formation

 Italy

A horseshoe crab belonging to the family Belinuridae. The type species is S. pontebbanus.

Tasmaniolimulus[196]

Gen. et sp. nov

Valid

Bicknell

Early Triassic[197]

Jackey Shale

 Australia

A member of Xiphosurida belonging to the family Austrolimulidae. The type species is T. patersoni.

Tuzoia lazizhaiensis[198]

Sp. nov

Valid

Wen et al.

Cambrian Stage 4

Balang Formation

 China

Zhiwenia[199]

Gen. et sp. nov

Valid

Du et al.

Cambrian Stage 3

Xiaoshiba Lagerstätte

 China

A non-trilobite artiopodan. Genus includes new species Z. coronata.

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