Broad-billed moa

Broad-billed moa; Temporal range: Pleistocene-Holocene
	Skull
Conservation status
	; Extinct (NZ TCS)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Aves
Infraclass:	Palaeognathae
Order:	†Dinornithiformes
Family:	†Emeidae
Genus:	†Euryapteryx; Haast, 1874
Species:	†E. curtus
Binomial name
	†Euryapteryx curtus; (Owen, 1846)
Synonyms
	List Cela Reichenbach 1853 non Moehring 1758 ; Celeus Bonaparte 1856 non Boie 1831 ; Zelornis Oliver 1949 ; Dinornis curtus Owen, 1846 ; Cela curtus (Owen 1846) Reichenbach, 1850 ; Celeus curtus (Owen 1846) Bonaparte, 1865 ; Anomalopteryx curta (Owen 1846) Lydekker 1891 ; Euryapteryx curtus (Owen 1846) Archey 1941 ; Mesopteryx species α Parker 1895 ; Euryapteryx exilis Hutton, 1897 ; Zelornis exilis (Hutton 1897) Oliver 1949 ; Euryapteryx tane Oliver 1949 ; Dinornis gravis Owen, 1870 ; Pachyornis gravis (Owen 1870) ; Euryapteryx pygmaeus Hutton 1891 non Pachyornis pygmaeus Hutton 1895 ; Emeus gravipes Lydekker, 1891 Euryapteryx gravipes (Lydekker 1891) Oliver 1930 ; Euryapteryx compacta Hutton 1893 ; Emeus crassus Parker 1895 non (Owen 1846) Reichenbach 1853 ; Euryapteryx ponderosa Hamilton 1898 non Hutton 1891 ; Emeus boothi Rothschild 1907 ; Emeus haasti Rothschild 1907 non Palaeocasuarius haasti Rothschild 1907 ; Zelornis haasti (Rothschild 1907) Oliver 1949 ; Euryapteryx haasti (Rothschild 1907) ; Emeus parkeri Rothschild 1907 ; Euryapteryx kuranui Oliver 1930 ; Euryapteryx geranoides Checklist Committee 1990 non Palapteryx geranoides ;

The broad-billed moa, stout-legged moa,^[5]^[6] or coastal moa (Euryapteryx curtus) is an extinct species of moa that was endemic to New Zealand.

Taxonomy

Euryapteryx curtus is a ratite and a member of the lesser moa family. The ratites are flightless birds with a sternum without a keel. They also have a distinctive palate.^[7]

A 2009 genetic study showed that the species Euryapteryx gravis is a junior synonym of E. curtus.^[8] Following this, a study published in 2010 explained size differences among the remains as being a result of sexual dimorphism.^[9] A 2012 morphological study interpreted them as subspecies instead.^[10] E. c. curtus, the nominate subspecies, was labeled the "coastal moa",^[1] while E. c. gravis was the "stout-legged moa".^[2]

The cladogram below follows a 2009 analysis by Bunce et al.:^[11]

Dinornithiformes

Megalapteryidae

	Megalapteryx didinus

Dinornithidae

Dinornis

	D. robustus

	D. novaezealandiae

Emeidae

Pachyornis

P. australis

	P. elephantopus

	P. geranoides

Anomalopteryx didiformis

	Emeus crassus

	Euryapteryx curtus

Habitat and distribution

Euryapteryx curtus was one of the most widespreas moa species, inhabiting mostly open areas.^[6] These moa lived on both the North and the South Islands of New Zealand, and remains have also been recovered from nearby Stewart Island. It inhabited lowland environments like duneland, forest, shrubland, and grassland.^[7]

Behaviour and ecology

As of 2006, half of all complete or mostly complete moa eggs in museum collections are likely broad-billed moa specimens.^[12] Of the specimens traditionally given the name Euryapteryx gravis, the eggs have an average length of 205 mm (8.1 in) and width of 143 mm (5.6 in), while the group traditionally assigned to the name Euryapteryx curtus had an average length of 122 mm (4.8 in) and width of 94 mm (3.7 in).^[12]

The species presumably went extinct for the same reasons as proposed for other moa: overhunting by the Māori (who called them "moa hakahaka") after their arrival on the islands in the 14th century.^[2]

References

^ ^a ^b "Euryapteryx curtus curtus. NZTCS". nztcs.org.nz. Retrieved 29 April 2023.
^ ^a ^b ^c "Euryapteryx curtus gravis. NZTCS". nztcs.org.nz. Retrieved 29 April 2023.
^ Owen, R. (1846). A History of British Fossil Mammals and Birds. London, UK: John Van Voorst.
^ Checklist Committee Ornithological Society of New Zealand (2010). "Checklist-of-Birds of New Zealand, Norfolk and Macquarie Islands and the Ross Dependency Antarctica" (PDF). Te Papa Press. Retrieved 4 January 2016.
^ "Stout-legged moa | New Zealand Birds Online".
^ ^a ^b Tennyson, Alan J. D. (2006). Extinct birds of New Zealand. Paul Martinson. Wellington, N.Z.: Te Papa Press. p. 36. ISBN 978-0-909010-21-8. OCLC 80016906.
^ ^a ^b Davies, S. J. J. F. (2003). "Moas". In Hutchins, Michael (ed.). Grzimek's Animal Life Encyclopedia. Vol. 8: Birds I: Tinamous and Ratites to Hoatzins (2nd ed.). Farmington Hills, MI: Gale Group. ISBN 0-7876-5784-0.
^ Bunce, Michael; Worthy, Trevor H.; Phillips, Matthew J.; Holdaway, Richard N.; Willerslev, Eske; Haile, James; Shapiro, Beth; Scofield, R. Paul; Drummond, Alexei; Kamp, Peter J. J.; Cooper, Alan (8 December 2009). "The evolutionary history of the extinct ratite moa and New Zealand Neogene paleogeography" (PDF). Proceedings of the National Academy of Sciences. 106 (49): 20646–20651. Bibcode:2009PNAS..10620646B. doi:10.1073/pnas.0906660106. PMC 2791642. PMID 19923428.
^ Gill, B. J. (2010). "Regional comparisons of the thickness of moa eggshell fragments (Aves: Dinornithiformes)". Records of the Australian Museum. 62: 115–122. doi:10.3853/j.0067-1975.62.2010.1535.
^ Worthy, T. H.; Scofield, R. P. (2012). "Twenty-first century advances in knowledge of the biology of moa (Aves: Dinornithiformes): a new morphological analysis and moa diagnoses revised". New Zealand Journal of Zoology. 39 (2): 87–153. doi:10.1080/03014223.2012.665060. S2CID 83768608.
^ Bunce et al. 2009.
^ ^a ^b Gill, B. J. (2006). "A Catalogue of Moa Eggs (Aves: Dinornithiformes)". Papahou: Records of the Auckland Museum. 43: 55–80. ISSN 1174-9202. JSTOR 42905885. Wikidata Q58623352.

External links

Unmasking the secrets of the extinct moa.

[NZTCS-1] "Euryapteryx curtus curtus. NZTCS". nztcs.org.nz. Retrieved 29 April 2023.

[NZTCS2-2] "Euryapteryx curtus gravis. NZTCS". nztcs.org.nz. Retrieved 29 April 2023.

[Brands-3] Owen, R. (1846). A History of British Fossil Mammals and Birds. London, UK: John Van Voorst.

[4] Checklist Committee Ornithological Society of New Zealand (2010). "Checklist-of-Birds of New Zealand, Norfolk and Macquarie Islands and the Ross Dependency Antarctica" (PDF). Te Papa Press. Retrieved 4 January 2016.

[5] "Stout-legged moa | New Zealand Birds Online".

[:0-6] Tennyson, Alan J. D. (2006). Extinct birds of New Zealand. Paul Martinson. Wellington, N.Z.: Te Papa Press. p. 36. ISBN 978-0-909010-21-8. OCLC 80016906.

[Davies-7] Davies, S. J. J. F. (2003). "Moas". In Hutchins, Michael (ed.). Grzimek's Animal Life Encyclopedia. Vol. 8: Birds I: Tinamous and Ratites to Hoatzins (2nd ed.). Farmington Hills, MI: Gale Group. ISBN 0-7876-5784-0.

[8] Bunce, Michael; Worthy, Trevor H.; Phillips, Matthew J.; Holdaway, Richard N.; Willerslev, Eske; Haile, James; Shapiro, Beth; Scofield, R. Paul; Drummond, Alexei; Kamp, Peter J. J.; Cooper, Alan (8 December 2009). "The evolutionary history of the extinct ratite moa and New Zealand Neogene paleogeography" (PDF). Proceedings of the National Academy of Sciences. 106 (49): 20646–20651. Bibcode:2009PNAS..10620646B. doi:10.1073/pnas.0906660106. PMC 2791642. PMID 19923428.

[9] Gill, B. J. (2010). "Regional comparisons of the thickness of moa eggshell fragments (Aves: Dinornithiformes)". Records of the Australian Museum. 62: 115–122. doi:10.3853/j.0067-1975.62.2010.1535.

[10] Worthy, T. H.; Scofield, R. P. (2012). "Twenty-first century advances in knowledge of the biology of moa (Aves: Dinornithiformes): a new morphological analysis and moa diagnoses revised". New Zealand Journal of Zoology. 39 (2): 87–153. doi:10.1080/03014223.2012.665060. S2CID 83768608.

[FOOTNOTEBunceWorthyPhillipsHoldaway2009-11] Bunce et al. 2009.

[Gill-12] Gill, B. J. (2006). "A Catalogue of Moa Eggs (Aves: Dinornithiformes)". Papahou: Records of the Auckland Museum. 43: 55–80. ISSN 1174-9202. JSTOR 42905885. Wikidata Q58623352.

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