Squalius

Squalius
	Squalius carolitertii
	Squalius pyrenaicus
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Order:	Cypriniformes
Family:	Leuciscidae
Subfamily:	Leuciscinae
Genus:	Squalius; Bonaparte, 1837
Type species
	Leuciscus squalus; Bonaparte, 1837
Synonyms
	Cephalopsis Fitzinger, 1873; Iberocypris Doadrio, 1980; Microlepis Bonaparte, 1846

Squalius is a genus of freshwater ray-finned fish belonging to the family Leuciscidae, the daces, Eurasian minnows and reklated fishes. The fishies in this genus are found in Europe and Asia.^[1] Hybridization is not rare in the Leuciscidae, including this genus. S. alburnoides is known to be of ancient hybrid origin, with the paternal lineage deriving from a prehistoric species related to Anaecypris; the latter mated with ancestral S. pyrenaicus. Present-day S. alburnoides mates with sympatric congeners of other species.^[2]

Species

These are the currently recognized species in this genus:^[3]

Squalius adanaensis Turan, Kottelat & Doğan, 2013 (Adana chub)
Squalius agdamicus S. N. Kamensky, 1901
Squalius alburnoides (Steindachner, 1866) (Calandino)
Squalius albus (Bonaparte, 1838) (Trasimeno chub)
Squalius anatolicus (Bogutskaya, 1997) (Beyşehir chub)
Squalius aradensis (Coelho, Bogutskaya, Rodrigues & Collares-Pereira, 1998) (Arade chub)
Squalius aristotelis Özuluğ & Freyhof, 2011 (Tuzla chub)
Squalius berak Heckel, 1843 (Mesopotamian chub)
Squalius caetobrigus Mendes, Perea, Sousa, Sousa-Santos & Doadrio, 2024
Squalius cappadocicus Özuluğ & Freyhof, 2011
Squalius carinus Özuluğ & Freyhof, 2011 (Chocolate chub)
Squalius carolitertii (Doadrio, 1988) (Northern Iberian chub)
Squalius castellanus Doadrio, Perea & Alonso, 2007 (Gallo chub)
Squalius cephalus (Linnaeus, 1758) (Common chub)
Squalius cii (J. Richardson, 1857) (Marmara chub)
Squalius fellowesii (Günther, 1868) (Aegean chub)
Squalius gaditanus Doadrio, Sousa-Santos & Perea, 2023
Squalius illyricus Heckel & Kner, 1857
Squalius irideus (Ladiges, 1960) (Anatolian ghizani)
Squalius keadicus (Stephanidis, 1971)
Squalius kosswigi (M. S. Karaman (sr), 1972)
Squalius kottelati Turan, Yılmaz & Kaya, 2009
Squalius laietanus Doadrio, Kottelat & de Sostoa, 2007 (Ebro chub)
Squalius latus Keyserling, 1861
Squalius lepidus Heckel, 1843 (Mesopotamian pike chub)
Squalius lucumonis (Bianco, 1983) (Etruscan chub)
Squalius malacitanus Doadrio & Carmona, 2006 (Malaga chub)
Squalius microlepis Heckel, 1843 (Imotski chub)
Squalius moreoticus (Stephanidis, 1971) (Stymphalia chub)
Squalius namak Khaefi, Esmaeili, Sayyadzadeh, Geiger & Freyhof, 2016
Squalius orientalis (Nordmann 1840)
Squalius orpheus Kottelat & Economidis, 2006 (Thracian chub)
Squalius palaciosi (Doadrio, 1980) (Bogardillo)
Squalius pamvoticus (Stephanidis, 1939) (Pamvotis chub)
Squalius peloponensis (Valenciennes, 1844) (Peloponnese chub)
Squalius platyceps Zupančič, Marić, Naseka & Bogutskaya, 2010
Squalius prespensis (Fowler, 1977) (Prespa chub)
Squalius pursakensis (Hankó (hu), 1925) (Sakarya chub)
Squalius pyrenaicus (Günther, 1868) (Southern Iberian chub)
Squalius recurvirostris Özuluğ & Freyhof, 2011 (Akşehir chub)
Squalius ruffoi (Bianco & Recchia, 1983)
Squalius semae Turan, Kottelat & Bayçelebi, 2017
Squalius seyhanensis Turan, Kottelat & Doğan, 2013 (Seyhan chub)
Squalius spurius Heckel, 1843 (Orontes chub)
Squalius squalus (Bonaparte 1837) (Italian chub)
Squalius svallize Heckel & Kner, 1857 (Neretva chub)
Squalius tartessicus Doadrio, Sousa-Santos & Perea, 2023
Squalius tenellus Heckel, 1843 (Livno chub)
Squalius torgalensis (Coelho, Bogutskaya, Rodrigues & Collares-Pereira, 1998) (Mira chub)
Squalius turcicus De Filippi, 1865 (Transcaucasian chub)
Squalius valentinus Doadrio & Carmona, 2006 (Eastern Iberian chub)
Squalius vardarensis S. L. Karaman, 1928 (Vardar chub)
Squalius verepi Turan, 2022
Squalius zrmanjae S. L. Karaman, 1928 (Zrmanja chub)

References

^ Froese, Rainer; Pauly, Daniel (eds.). "Species in genus Squalius". FishBase. October 2016 version.
^ Collares-Pereira, M.J. & Coelho, M.M. (2010): Reconfirming the hybrid origin and generic status of the Iberian cyprinid complex Squalius alburnoides. Journal of Fish Biology, 76 (3): 707–715.
^ Eschmeyer, William N.; Fricke, Ron & van der Laan, Richard (eds.). "Species in the genus Squalius". Catalog of Fishes. California Academy of Sciences. Retrieved 5 May 2025.

[FB-1] Froese, Rainer; Pauly, Daniel (eds.). "Species in genus Squalius". FishBase. October 2016 version.

[Collares2010-2] Collares-Pereira, M.J. & Coelho, M.M. (2010): Reconfirming the hybrid origin and generic status of the Iberian cyprinid complex Squalius alburnoides. Journal of Fish Biology, 76 (3): 707–715.

[Cof_genus-3] Eschmeyer, William N.; Fricke, Ron & van der Laan, Richard (eds.). "Species in the genus Squalius". Catalog of Fishes. California Academy of Sciences. Retrieved 5 May 2025.

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